Mites associated with bees of the World, species list

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Species and subspecies of mites associated with bees of the World

(habitat generalists or unspecific taxa repetitively found in association with honeybees, bumblebees, or stingless bees are also included)

Apis= associated with Apis spp., Bombus=associated with Bombus spp.

(under construction)

ACARIFORMES Zachvatkin, 1952

Sarcoptiformes Reuter, 1909
- Astigmata Canestrini, 1891
  - Family Acaridae Latreille, 1802
    - Genus 'Acotyledon'
      - 'Acotyledon' palaeorhizae Samšiňák, 1968
    - Genus Acarus
      - Acarus farris (Oudemans, 1905) [Apis]
      - Acarus gracilis Hughes, 1957 [Apis]
      - Acarus immobilis Griffiths, 1964 [Apis]
      - Acarus siro Linnaeus, 1758 [Apis Bombus]
      - Acarus tyrophagoides Zachvatkin, 1941 [Apis]
    - Genus Acotyledon
      - Acotyledon paradoxa Oudemans, 1903 [Apis]
    - Genus Aleuroglyphus
      - Aleuroglyphus ovatus (Troupeau, 1878) [Apis]
      - Aleuroglyphus siculus (Fumouze and Robin, 1867) [Apis]
    - Genus Cerophagopsis
      - Cerophagopsis furcata Fain, 1974
      - Cerophagopsis heriades (Fain and Pauly, 2001)
      - Cerophagopsis skorikovi Zachvatkin, 1941
      - Cerophagopsis trigona Fain and Heard, 1987
    - Genus Cosmoglyphus
      - Cosmoglyphus oudemansi (Zachvatkin, 1937) [Apis]
      - Cosmoglyphus sp. Haragsim at al., 1978 [Apis]
    - Genus Diadasiopus
      - Diadasiopus alexanderi OConnor and Daneshvar, 1999
      - Diadasiopus eickworti OConnor, 1997
    - Genus Forcellinia
      - Forcellinia faini Delfinado-Baker and Baker, 1989 [Apis]
      - Forcellinia galleriella Womersley, 1963 [Apis]
      - Forcellinia sp. n.
    - Genus Halictacarus
      - Halictacarus halicti Mahunka, 1975
      - Halictacarus halicti (Putatunda and Kapil, 1995)
      - Halictacarus indicus (Putatunda and Kapil, 1995)
      - Halictacarus orientalis (Putatunda and Kapil, 1995)
      - Halictacarus spp. Putatunda&Arbol2003
      - Halictacarus zakhvatkini (Delfinado and Baker, 1976)
    - Genus Histiogaster
      - Histiogaster bacchus Zachvatkin, 1941 [Apis]
    - Genus Horstia
      - Horstia allaeri Fain, 1974
      - Horstia amplisucta Fain and Camerik, 1978 [Bombus]
      - Horstia brasiliensis Fain and Camerik, 1978 [Bombus]
      - Horstia glabra (Vitzthum, 1920) (=Horstia scutata Fain,1974)
      - Horstia gibba (Oudemans, 1924)
      - Horstia helenae (Oudemans, 1902)
      - Horstia major Alzuet and Abrahamovich, 1987
      - Horstia malaysiensis (Fain, Lukoschus and Nadchatram, 1982)
      - Horstia minor Alzuet and Abrahamovich, 1987
      - Horstia monstruosa (Vitzthum, 1920)
      - Horstia ornata (Oudemans, 1900)
      - Horstia pulcherrima (Vitzthum, 1912)
      - Horstia rwandae Fain, 1984
      - Horstia splendidissima Vitzthum, 1941
      - Horstia tanzaniensis Fain, 1974
      - Horstia trifilis Canestrini, 1920
      - Horstia virginica Baker, 1962
    - Genus Horstiella
      - Horstiella armata Turk, 1948
      - Horstiella concentrica Ochoa and OConnor, 2000
      - Horstiella megamyzidos Ochoa and OConnor, 2000
      - Horstiella mourei Ochoa and OConnor, 2000
      - Horstiella quadrata Ochoa and OConnor, 2000
      - Horstiella snellingi Ochoa and OConnor, 2000
      - Horstiella variabilis Ochoa and OConnor, 2000
    - Genus Kuzinia
      - Kuzinia affinis Delfinado and Baker, 1976 [Bombus]
      - Kuzinia americana Delfinado and Baker 1976 [Bombus]
      - Kuzinia dispar Delfinado and Baker, 1976 [Bombus]
      - Kuzinia evae Putatunda, Aggarwal and Kapil, 1984 [Bombus]
      - Kuzinia laevis (Dujardin, 1849) [Apis Bombus]
      - Kuzinia nova Volgin and Kadzhaya, 1973 [Bombus]
      - Kuzinia spp. Kumar1996; Putatunda&Arbol2003; Husband, 1994 [Apis Bombus]
      - Kuzinia woykei Putatunda, Aggarwal and Kapil, 1984 [Bombus]
    - Genus Lasioacarus
      - Lasioacarus morsei (El-Banhawy and Abou-Awad, 1990) [Apis]
      - Lasioacarus nidicolus Kadzhaya and Sevastyanov, 1967 [Apis]
    - Genus Medeus
      - Medeus ithacaensis OConnor, 1996
    - Genus Megachilopus
      - Megachilopus uellensis Fain, 1974
    - Genus Neoacotyledon
      - Neoacotyledon rhizoglyphoides (Zakhvatkin, 1937) [Apis]
    - Genus Neohorstia
      - Neohorstia mamillata Zachvatkin, 1941
    - Genus Rhizoglyphus
      - Rhizoglyphus robini Claparède, 1869 [Apis]
      - Rhizoglyphus sp. Eickwort in Fain & al. 1999; Kamali et al., 2001 [Apis]
    - Genus Sancassania
      - Sancassania (Ctenocolletacarus) brevirostris Fain, 1984
      - Sancassania (Ctenocolletacarus) grandior Fain, 1984
      - Sancassania (Ctenocolletacarus) longirostris Fain, 1984
      - Sancassania boharti (Cross, 1968)
      - Sancassania rodionovi (Zachvatkin, 1935) (=Sancassania berlesei auct. non (Michael, 1903)) [Apis]
      - Sancassania sphaerogaster Zachvatkin, 1937 (=Caloglyphus michaeli (поп Oudemans, 1924); Caloglyphus mycophagus auct. (non. Mégnin, 1874); Sancassania mycophaga auct. non Mégnin, 1874) [Apis]
      - Sancassania spp. Malabanan&Corpuz-Raros, 1988; Eickwort, 1979 [Apis]
    - Genus Schulzea
      - Schulzea caucasica Zakhvatkin, 1941
      - Schulzea pamirensis Zachvatkin, 1941
    - Genus Schwiebea
      - Schwiebea tshernyshevi Zachvatkin, 1941 [Apis]
    - Genus Sennertionyx
      - Sennertionyx manicati (Giard, 1900)
    - Genus Thectochloracarus
      - Thectochloracarus neotropicalis Fain, Engel, Flechtmann, OConnor, 1999
    - Genus Thyreophagus
      - Thyreophagus corticalis (Michael, 1885) [Apis]
      - Thyreophagus entomophagus (Laboulbène, 1852) [Apis Bombus]
      - Thyreophagus odyneri Fain, 1982 [Apis]
    - Genus Tyroborus
      - Tyroborus sp. Grobov, 1975 [Apis]
    - Genus Tyrolichus
      - Tyrolichus casei Oudemans, 1910 [Apis]
      - Tyrolichus sp. Kamali at al., 2001;Mosaddegh, 1997 [Apis]
    - Genus Tyrophagus
      - Tyrophagus curvipenis Fain and Fauvel, 1993 [Apis]
      - Tyrophagus debrivorus Chinniah and Mohanasundaram, 1996 [Apis]
      - Tyrophagus longior (Gervais, 1844) [Apis]
      - Tyrophagus mixtus Volgin, 1948 [Apis]
      - Tyrophagus perniciosus Zakhvatkin, 1941 [Apis]
      - Tyrophagus putrescentiae (Schrank, 1781) [Apis Bombus]
      - Tyrophagus savasi Lynch, 1989 [Apis]
      - Tyrophagus similis Volgin, 1949 [Apis Bombus]
      - Tyrophagus tropicus Robertson, 1959 [Apis]
      - Tyrophagus vanheurni Oudemans, 1924 [Apis]
  - Family Aeroglyphidae Zachvatkin, 1941
    - Genus Aeroglyphus
      - Aeroglyphus peregrinans (Berlese, 1892) [Apis Bombus]
      - Aeroglyphus robustus (Banks, 1906) [Apis]
    - Genus Glycycometus
      - Glycycometus combus (Chinniah and Mohanasundaram, 1996) [Apis]
      - Glycycometus geniculatus (Vitzthum, 1920)
      - Glycycometus malaysiensis (Fain and Nadchatram, 1980) [Apis]
      - Glycycometus molitor (Volgin and Akimov, 1975) (=Austroglycyphagus geniculatus auct. non (Vitzthum, 1919); Glycycometus hughesae (Fain, 1976)) [Apis]
      - Glycycometus rwandae (Fain, 1976) [Apis]
      - Glycycometus thailandicus (Fain and Gerson, 1990) [Apis]
      - Glycycometus 'orientalis' nom. nud. [Apis]
  - Family Astigmata fam.
    - Genus Astigmata
      - Astigmata gen. sp. Engel, 1996
  - Family Carpoglyphidae Oudemans, 1923
    - Genus Carpoglyphus
      - Carpoglyphus lactis (Linnaeus, 1758) (=Acarus passularum Hering) [Apis]
      - Carpoglyphus munroi Hughes, 1952 [Apis]
  - Family Chaetodactylidae Zachvatkin, 1941
    - Genus Achaetodactylus
    - Genus Centriacarus
      - Centriacarus guahibo Klimov and OConnor, 2007
      - Centriacarus turbator Klimov and OConnor, 2007
    - Genus Chaetodactylus
      - Chaetodactylus abditus Klimov and OConnor, 2004
      - Chaetodactylus anthidii (Oudemans, 1911)
      - Chaetodactylus antillarum Klimov and OConnor, 2008
      - Chaetodactylus azteca Klimov and OConnor, 2008
      - Chaetodactylus birulai Zachvatkin, 1941 (=Chaetodactylus poetae Samšiňák, 1973)
      - Chaetodactylus chrysidis Fain and Baugnée, 1996 (=Chaetodactylus chrysidis aurulenticola Fain and Baugnée, 1996)
      - Chaetodactylus claudus Klimov and OConnor, 2008
      - Chaetodactylus claviger Oudemans, 1924 [Apis]
      - Chaetodactylus dalyi (Fain, 1974)
      - Chaetodactylus dementjevi Zachvatkin, 1941
      - Chaetodactylus furunculus Klimov and OConnor, 2008
      - Chaetodactylus gibbosi Klimov and OConnor, 2004
      - Chaetodactylus hirashimai Kurosa, 1987
      - Chaetodactylus hopliti Klimov and OConnor, 2008
      - Chaetodactylus kouboy Klimov and OConnor, 2008
      - Chaetodactylus krombeini Baker, 1962
      - Chaetodactylus lassulus Klimov and OConnor, 2007
      - Chaetodactylus lithurgi Klimov and OConnor, 2004
      - Chaetodactylus ludwigi (Trouessart, 1904)
      - Chaetodactylus melitomae Klimov and OConnor, 2007
      - Chaetodactylus micheneri Klimov and OConnor, 2008
      - Chaetodactylus nipponicus Kurosa, 1987
      - Chaetodactylus osmiae (Dufour, 1839) (=Chaetodactylus mahunkai Samšiňák, 1973) [Apis]
      - Chaetodactylus reaumuri (Oudemans, 1905)
      - Chaetodactylus rozeni Klimov and OConnor, 2008
      - Chaetodactylus zachvatkini Klimov and OConnor, 2008
    - Genus Roubikia
    - Genus Sennertia
      - Sennertia aldeodadi Haitlinger, 2000
      - Sennertia alfkeni (Oudemans, 1900) (=Sennertia japonica (Oudemans, 1900))
      - Sennertia americana Delfinado and Baker, 1976
      - Sennertia antarctica (Trägårdh, 1907)
      - Sennertia argentina Vitzthum, 1941
      - Sennertia augustii Alzuet and Abrahamovich, 1990
      - Sennertia basilewskyi Fain, 1974
      - Sennertia benoiti Fain, 1974
      - Sennertia bifida Kurosa, 2003
      - Sennertia bifilis (Canestrini, 1897)
      - Sennertia caffra Vitzthum, 1919
      - Sennertia cantabrica Zachvatkin, 1941
      - Sennertia capensis Fain, 1971
      - Sennertia cerambycina (Scopoli, 1763)
      - Sennertia ceratinarum Fain, 1974
      - Sennertia congoicola Fain, 1971
      - Sennertia dalyi Fain, 1980
      - Sennertia dalyi nilotica Fain, 1980
      - Sennertia delfinadoae Fain, 1981 (=Sennertia bakeri Ramaraju and Mohanasundaram, 2001)
      - Sennertia devincta Klimov and OConnor, 2007
      - Sennertia dissimilis Zachvatkin, 1941
      - Sennertia donaldi Turk, 1948
      - Sennertia duweinii Sherbef and Duweini, 1980
      - Sennertia egyptiaca Elbadry, 1971
      - Sennertia elseni Fain, 1971
      - Sennertia faini Baker and Delfinado-Baker, 1983 [Apis]
      - Sennertia flabellifera Oudemans, 1924
      - Sennertia frontalis Vitzthum, 1941
      - Sennertia gargantua Zachvatkin, 1941
      - Sennertia greeni (Oudemans, 1917)
      - Sennertia haustrifera Klimov and OConnor, 2008
      - Sennertia herminae Haitlinger, 1999
      - Sennertia hipposideros (Oudemans, 1902)
      - Sennertia horrida (Vitzthum, 1912) (=Sennertia sumatrensis Oudemans, 1924; Sennertia leucothorae Ramaraju and Mohanasundaram, 2001)
      - Sennertia hurdi Klimov and OConnor, 2008
      - Sennertia ignota Delfinado and Baker, 1976
      - Sennertia indica Delfinado and Baker, 1976
      - Sennertia jeanalexi Fain, 1971
      - Sennertia koptorthosomae (Oudemans, 1901)
      - Sennertia latipilis Fain, 1974
      - Sennertia lauta Klimov and OConnor, 2007
      - Sennertia leclercqi Fain, 1971
      - Sennertia leei Fain, 1982
      - Sennertia longipilis Alzuet and Abrahamovich, 1987
      - Sennertia loricata Klimov and OConnor, 2008
      - Sennertia lucrosa Klimov and OConnor, 2008
      - Sennertia madagascarensis Fain, 1971
      - Sennertia mesotrichia Fain, 1971
      - Sennertia micheli Fain, 1971
      - Sennertia monicae Fain, 1971
      - Sennertia morstatti Vitzthum 1914 (=Sennertia moandensis Fain, 1971)
      - Sennertia oudemansi Zachvatkin, 1941
      - Sennertia perturbans Vitzthum, 1919
      - Sennertia pirata Klimov and OConnor, 2008
      - Sennertia potanini Zachvatkin, 1941
      - Sennertia queenslandica Womersley, 1941
      - Sennertia ratiocinator Klimov and OConnor, 2007
      - Sennertia recondita Klimov and OConnor, 2008
      - Sennertia robusta Delfinado and Baker, 1976 (=Sennertia carpenteri Ramaraju and Mohanasundaram, 2001)
      - Sennertia roepkei Oudemans, 1924
      - Sennertia sayutara Klimov and OConnor, 2007
      - Sennertia scutata Fain, 1974
      - Sennertia segnis Klimov and OConnor, 2008
      - Sennertia shimanukii Baker and Delfinado-Baker, 1983 [Apis]
      - Sennertia simplex (Trägårdh, 1904)
      - Sennertia sodalis Klimov and OConnor, 2008
      - Sennertia spinifera Fain, 1974
      - Sennertia splendidulae Alzuet and Abrahamovich, 1989
      - Sennertia spp. Kumar 1996; Putatunda and Arbol, 2003 [Apis Bombus]
      - Sennertia surinamensis Fain and Lukoschus, 1971
      - Sennertia tanythrix Fain, 1971
      - Sennertia tunisiana Fain, 1980
      - Sennertia vaga Klimov and OConnor, 2008
      - Sennertia vanderhammeni brevipilis Fain, 1974
      - Sennertia vanderhammeni Fain, 1974
      - Sennertia varicosa Fain, 1971
      - Sennertia vitzthumi Fain, 1981
      - Sennertia zhelochovtsevi Zachvatkin, 1941
  - Family Chortoglyphidae Berlese, 1897
    - Genus Chortoglyphus
      - Chortoglyphus arcuatus (Troupeau, 1879) [Apis]
  - Family Echimyopodidae Fain, 1967
    - Genus Blomia
      - Blomia nr. freemani Hughes, 1948 [Apis]
  - Family Gaudiellidae Atyeo, Baker & Delfinado, 1974
    - Genus Cerophagus
      - Cerophagus granulatus (Dujardin, 1849) (=Cerophagus gracilis Vitzthum, 1912; Glycyphagus bomborum Oudemans, 1902) [Bombus]
      - Cerophagus nearcticus OConnor, 1992 [Bombus]
    - Genus Gaudiella
      - Gaudiella camargoi (Fain and Rosa, 1983) (=Trigonacoptes camargoi Fain and Rosa, 1983)
      - Gaudiella minuta Atyeo, Baker and Delfinado, 1974
    - Genus Meliponopus
      - Meliponopus palpifer Fain and Flechtmann, 1985
    - Genus Partamonacoptes
      - Partamonacoptes brasiliensis Fain and Rosa, 1983
    - Genus Platyglyphus
      - Platyglyphus malayanus Kurosa, 1976
  - Family Glycyphagidae Berlese, 1897
    - Genus Ctenoglyphus
      - Ctenoglyphus plumiger (C. L. Koch, 1835) [Apis]
    - Genus Diamesoglyphus
      - Diamesoglyphus intermedius (G. Canestrini, 1888) [Apis]
    - Genus Glycyphagus
      - Glycyphagus (Glycyphagus) domesticus (De Geer, 1778) [Apis]
      - Glycyphagus (Glycyphagus) ornatus Kramer, 1881 [Apis Bombus]
      - Glycyphagus (Lepidoglyphus) privatus Oudemans, 1903 [Apis]
      - Glycyphagus (Lepidoglyphus) destructor (Schrank, 1781) [Apis Bombus]
      - Glycyphagus (Lepidoglyphus) fustifer (Oudemans, 1903) [Apis]
      - Glycyphagus (Lepidoglyphus) michaeli (Oudemans, 1903) [Apis]
    - Genus Gohieria
      - Gohieria fusca (Oudemans, 1902) [Apis]
    - Genus Lophuromyopus
      - Lophuromyopus sp. Kamali at al., 2001 [Apis]
    - Genus Xenoryctes
      - Xenoryctes krameri (Michael, 1886) [Bombus]
  - Family Hemisarcoptidae Oudemans, 1904
    - Genus 'Nanacarus'
      - 'Nanacarus' nominis Woodring, 1966
    - Genus Nanacarus
      - Nanacarus minutus (Oudemans, 1901)
  - Family Heterocoptidae Fain, 1967
    - Genus Erotylocoptes
      - Erotylocoptes anthophora Fain, 1987
  - Family Histiostomatidae Berlese, 1897
    - Genus Anoetus
      - Anoetus alicola (Dujardin, 1849) [Apis]
      - Anoetus debilis Woodring, 1973
      - Anoetus eickworti (Woodring, 1973)
      - Anoetus halicticola (Fain and Erteld, 1998)
      - Anoetus halictonida (Woodring, 1973)
      - Anoetus kaszabi Mahunka, 1969
      - Anoetus ligulotrichus Mahunka, 1974
      - Anoetus orientalis Mahunka, 1974
      - Anoetus spp. Eickwort, 1979; Dolphin, 1979 (Histiostoma); De Jong et al., 1985 [Apis]
      - Anoetus szelenyii Mahunka, 1974
      - Anoetus tunisiensis Mahunka, 1974
      - Anoetus vexarus Woodring, 1973
      - Anoetus sp. Walter et al., 2002
    - Genus Glyphanoetus
      - Glyphanoetus nomiensis Cross, 1968
    - Genus Histiostoma
      - Histiostoma conclavicola (Oudemans, 1929)
      - Histiostoma feroniarum (Dufour, 1839) [Apis]
      - Histiostoma heriades Fain and Pauly, 2001
      - Histiostoma inquilinus Cross, 1968
      - Histiostoma polypori (Oudemans, 1914) [Apis]
    - Genus Myianoetus
      - Myianoetus setipes (Koch, 1847) [Apis]
  - Family Meliponocoptidae Fain & Rosa, 1983
    - Genus Meliponocoptes
      - Meliponocoptes nidicolus Fain and Rosa, 1983
      - Meliponocoptes orphanus Fain and Rosa, 1983
      - Meliponocoptes scutatus Fain and Rosa, 1983
    - Genus Meliponoecius
      - Meliponoecius flechtmanni Fain and Rosa, 1983
  - Family Pyroglyphidae Cunliffe, 1958
    - Genus Dermatophagoides
      - Dermatophagoides farinae Hughes, 1961 [Apis]
      - Dermatophagoides pteronyssinus (Trouessart, 1897) [Apis]
      - Dermatophagoides sp. Kumar, 1996 [Apis]
  - Family Suidasiidae Hughes, 1948
    - Genus Suidasia
      - Suidasia pontifica Oudemans, 1905 [Apis]
    - Genus Tortonia
      - Tortonia dogaressa Zakhvatkin, 1941
      - Tortonia fluctuata Oudemans, 1917 [Bombus]
      - Tortonia intermedia (Oudemans, 1902)
      - Tortonia quadridens Baker, 1962
      - Tortonia smitsvanburgsti Oudemans, 1911
      - Tortonia sp. Kuhlmann, 1998
      - Tortonia volfi (Samšiňák, 1973)
  - Family Winterschmidtiidae Oudemans, 1923
    - Genus Saproglyphus
      - Saproglyphus hagensis (Oudemans, 1911)
      - Saproglyphus reticulatus Zachvatkin, 1941
      - Saproglyphus sp. Haragsim at al., 1978 [Apis]
    - Genus Vidia
      - Vidia bomborum (Leonardi, 1900) [Bombus]
      - Vidia hirsuta OConnor and Eickwort, 1988
      - Vidia latimanus OConnor and Eickwort, 1988
      - Vidia lineata Oudemans, 1917
      - Vidia rubi OConnor and Eickwort, 1988
      - Vidia texana OConnor and Eickwort, 1988
      - Vidia undulata Oudemans, 1905
      - Vidia sp. Putatunda and Arbol, 2003 [Apis]
      - Vidia sp1 Sumangala et al., 2001 [Apis]
      - Vidia sp2 Sumangala et al., 2001 [Apis]
      - Vidia texana OConnor and Eickwort, 1988
      - Vidia undulata Oudemans, 1905
    - Genus Winterschmidtiidae
      - Winterschmidtiidae gen. n. Eickwort, 1994
- Oribatida van der Hammen, 1968
  - Family Carabodidae C.L.Koch, 1837
    - Genus Carabodes
      - Carabodes minusculus Berlese, 1923 [Apis]
  - Family Ceratozetidae Jacot, 1925
    - Genus Ceratozetes
      - Ceratozetes sp. nr. longispina Jacot, 1936 [Bombus]
    - Genus Diapterobates
      - Diapterobates humeralis (Hermann, 1804) [Apis]
    - Genus Trichoribates
      - Trichoribates sp. Crozier, 1989 [Apis]
      - Trichoribates trimaculatus (C.L. Koch, 1835) [Apis]
  - Family Crotoniidae Thorell, 1876
    - Genus Camisia
      - Camisia spinifer Koch, 1836 [Apis]
    - Genus Platynothrus
      - Platynothrus peltifer (C.L. Koch, 1839) [Apis]
  - Family Damaeidae Berlese, 1896
    - Genus Belba
      - Belba sp. Chmielewski, 1971 [Bombus]
    - Genus Epidamaeus
      - Epidamaeus kamaensis (Sellnick, 1926) [Apis]
      - Epidamaeus sp. Grobov, 1978 [Apis]
  - Family Eulohmanniidae Grandjean, 1931
    - Genus Lohmannia
      - Lohmannia (Eulohmannia) ribagai Berlese, 1910
  - Family Euzetidae Grandjean, 1954
    - Genus Euzetidae
      - Euzetidae gen. sp. Grobov, 1978 [Apis]
  - Family Galumnidae Jacot, 1925
    - Genus Galumna
      - Galumna microfissum Hammer, 1968 [Bombus]
      - Galumna sp. Piletskaya and Zaloznaya, 2004 [Apis]
  - Family Haplochthoniidae van der Hammen, 1959
    - Genus Amnemochthonius
      - Amnemochthonius taeniophorus Grandjean, 1948 [Apis]
  - Family Haplozetidae Grandjean, 1936
    - Genus Peloribates
      - Peloribates europaeus Willmann, 1935 [Apis]
      - Peloribates rangiroaensis Hammer, 1972 [Apis]
  - Family Hypochthoniidae Berlese, 1910
    - Genus Eohypochthonius
      - Eohypochthonius vermicularis Hammer, 1979 [Apis]
  - Family Liacaridae Sellnick, 1928
    - Genus Birsteinius
      - Birsteinius clavatus Krivolutsky, 1965 [Apis]
  - Family Malaconothridae Berlese, 1916
    - Genus Trimalaconothrus
      - Trimalaconothrus foveolatus Willmann, 1931 [Apis]
  - Family Micreremidae Grandjean, 1954
    - Genus Micreremus
      - Micreremus brevipes (Michael, 1888) [Apis]
  - Family Mochlozetidae Grandjean, 1960
    - Genus Pelokylla
      - Pelokylla malabarica Adolph and Haq, 1982 [Apis]
  - Family Mycobatidae Grandjean, 1954
    - Genus Minguezetes
      - Minguezetes hexagonus (Berlese, 1908) [Apis]
    - Genus Punctoribates
      - Punctoribates sp. Grobov, 1978 [Apis]
  - Family Oppiidae Grandjean, 1951
    - Genus Oppia
      - Oppia concolor (C.L. Koch, 1844) [Apis]
      - Oppia nitens Koch, 1836 [Apis]
      - Oppia unicarinata (Paoli, 1908) [Apis]
  - Family Oribatulidae Thor, 1929
    - Genus Paraleius
      - Paraleius leontonychus (Berlese, 1910) [Apis]
    - Genus Zygoribatula
      - Zygoribatula frisiae (Oudemans, 1900) [Apis]
      - Zygoribatula propinqua (Oudemans, 1902) [Apis]
      - Zygoribatula spp. Grobov, 1975; Kamali at al., 2001 [Apis]
      - Zygoribatula undulata Berlese, 1916 [Apis]
  - Family Pediculochelidae Lavoipierre, 1946
    - Genus Paralycus
      - Paralycus raulti (Lavoipierre, 1947) [Apis]
  - Family Phthiracaridae Perty, 1841
    - Genus Phthiracarus
      - Phthiracarus italicus (Oudemans, 1900) [Apis]
  - Family Scheloribatidae Grandjean, 1933
    - Genus Corynobates
      - Corynobates clavisetus (Mahunka, 1988) [Apis]
    - Genus Scheloribates
      - Scheloribates labyrinthicus Jeleva 1962 [Apis]
      - Scheloribates laevigatus (C.L. Koch, 1836) [Apis]
      - Scheloribates latipes (C. L. Koch, 1844) [Apis]
      - Scheloribates pallidulus (C.L. Koch, 1841) [Apis]
      - Scheloribates sp. Crozier, 1989 [Apis]
      - Scheloribates zealandicus Hammer, 1967 [Bombus]
  - Family Tegoribatidae Grandjean, 1954
    - Genus Tegoribates
      - Tegoribates latirostris (C.L. Koch,1844) [Apis]
  - Family Trhypochthoniidae Willmann, 1931
    - Genus Trhypochthonius
      - Trhypochthonius tectorum (Berlese, 1896) [Apis]
Trombidiformes Reuter, 19091
- Prostigmata Kramer, 1877
  - Family Anystidae Oudemans, 1936
    - Genus Anystis
      - Anystis spp. De Jong et al., 1985; Grobov, 1975,1978 [Apis]
    - Genus Erythracarus
      - Erythracarus parietinus (Hermann, 1804) [Apis]
  - Family Bdellidae Dugès, 1834
    - Genus Bdella
      - Bdella iconica Berlese [Bombus]
      - Bdella sp. Banaszak, 1980 [Apis]
    - Genus Bdellidae
      - Bdellidae spp. Chmielewski, 1991; Tareev,1972 [Apis]
    - Genus Bdellodes
      - Bdellodes lapidaria (Kramer, 1881) [Apis]
  - Family Caligonellidae Grandjean, 1944
    - Genus Neognathus
      - Neognathus nr. eupalopus Meyer and Ueckermann, 1989 [Apis]
  - Family Camerobiidae Southcott, 1957
    - Genus Neophyllobius
      - Neophyllobius sp. Kamali at al., 2001; Mosaddegh, 1997 [Apis]
  - Family Cheyletidae Leach, 1815
    - Genus Acaropsella
      - Acaropsella sp. Haragsim at al., 1977 [Apis]
    - Genus Acaropsellina
      - Acaropsellina sollers (Kuzin, 1940) [Apis]
    - Genus Bak
      - Bak furcatus Gerson and Fain, 1991 [Apis]
    - Genus Cheletomimus
      - Cheletomimus bregetovae (Volgin, 1969) [Apis]
    - Genus Cheletomorpha
      - Cheletomorpha lepidopterorum (Shaw, 1794) [Apis]
    - Genus Cheletophyes
      - Cheletophyes apicola Fain, Lukoschus and Nadchatram, 1980
      - Cheletophyes clavipilis Fain, Lukoschus and Nadchatram, 1980
      - Cheletophyes decorus Bochkov and Klimov, 2004
      - Cheletophyes eckerti Summers and Price, 1970
      - Cheletophyes indiacus Smiley and Whitaker, 1981 (=Cheletophyes xylocopae Ramaraju et Mohanasundaram, 1999)
      - Cheletophyes mbomba Bochkov, Klimov and OConnor, 2006
      - Cheletophyes occisor Bochkov and Klimov, 2004
      - Cheletophyes panamensis Klompen, Méndez and Lukoschus, 1984
      - Cheletophyes sp. Walter at al., 2002
      - Cheletophyes torridae Bochkov, Klimov and OConnor, 2006
      - Cheletophyes venator (Vitzthum, 1920) (=Cheletophyes aurorae Haitlinger 2000)
      - Cheletophyes vespae Fain and Bochkov, 2001
      - Cheletophyes vitzthumi Oudemans, 1914
    - Genus Chelacheles
      - Chelacheles temoak Bochkov and OConnor, 2004
    - Genus Cheyletus
      - Cheyletus carnifex Zachvatkin, 1935 [Apis]
      - Cheyletus eruditus Schrank, 1781 [Apis Bombus]
      - Cheyletus malaccensis Oudemans, 1903 [Apis]
    - Genus Eucheyletia
      - Eucheyletia bakeri Volgin, 1963 [Bombus]
      - Eucheyletia flabellifera (Michael, 1878) [Apis]
    - Genus Nodele
      - Nodele (Aztecocheyletus) conquistador Bochkov and Klimov, 2004
  - Family Cunaxidae Thor, 1902
    - Genus Armascirus
      - Armascirus taurus (Kramer, 1881) [Apis]
    - Genus Cunaxa
      - Cunaxa lukoschusi Smiley, 1992 [Apis]
    - Genus Pulaeus
      - Pulaeus rimandoi Corpuz-Raros, 1996 [Apis]
  - Family Ereynetidae Oudemans, 1931
    - Genus Ereynetes
      - Ereynetes (Anereynetes) meliponae Flechtmann, Fain, and Leal, 1985
      - Ereynetes (Ereynetes) boharti Hunter and Cross, 1968
  - Family Eriophyidae Nalepa, 1898
    - Genus Cecidophyopsis
      - Cecidophyopsis ribis (Westwood, 1869) [Apis]
    - Genus Eriophyes
      - Eriophyes litchii Keifer, 1943 [Apis]
      - Eriophyes tiliae (Pagenstecher, 1857) [Apis]
  - Family Erythraeidae Robineau-Desvoidy, 1828
    - Genus Erythraeidae
      - Erythraeidae sp. Malabanan and Corpuz-Raros, 1998 [Apis]
    - Genus Leptus
      - Leptus ariel Southcott, 1989 [Apis]
      - Leptus monteithi Southcott, 1993
      - Leptus sp. Wilson, Rubink & Collins, 1990 [Apis]
  - Family Neotrombidiidae Feider, 1959
    - Genus Neotrombidium
      - Neotrombidium sp. Klimov (unpubl.)
  - Family Podapolipidae Ewing, 1922
    - Genus Locustacarus
      - Locustacarus buchneri (Stammer, 1951) [Apis Bombus]
  - Family Pyemotidae Oudemans, 1937
    - Genus Pyemotes
  - Family Neopygmephoridae Cross, 1965
    - Genus Kerdabania
      - Kerdabania quadrata (Ewing, 1917) [Apis]
    - Genus Insensilla
      - Insensilla duplicis Kurosa, 2009
      - Insensilla lasioglossi Kurosa, 2009
    - Genus Bakerdania
      - Bakerdania amplus (Krczal, 1959)
    - Genus Parapygmephorus
      - Parapygmephorus (Parapygmephorus) natalensis Cross, 1965
      - Parapygmephorus (Parapygmephorus) spp.
      - Parapygmephorus (Sicilipes) costaricanus Rack and Eickwort, 1980
      - Parapygmephorus (Sicilipes) halictinis Cross, 1965
      - Parapygmephorus (Sicilipes) sp2 Cross, 1965
      - Parapygmephorus (Parapygmephorus) aestivus (Berlese, 1886)
      - Parapygmephorus (Parapygmephorus) crossi Mahunka, 1974
      - Parapygmephorus (Parapygmephorus) nr. crossi Mahunka, 1974
      - Parapygmephorus (Parapygmephorus) delyorum Mahunka, 1980
      - Parapygmephorus (Parapygmephorus) khorasanicus Hajiqanbar and Khaustov, 2011
      - Parapygmephorus (Parapygmephorus) luxtoni (Mahunka, 1970)
      - Parapygmephorus (Parapygmephorus) magnisetosus Khaustov and Zaloznaya, 2011
      - Parapygmephorus (Parapygmephorus) pappi Mahunka, 1974
      - Parapygmephorus sp1 Eickwort & Eickwort, 1971
      - Parapygmephorus sp2 Mohamed & Soliman, 1974
  - Family Pygmephoridae Cross, 1965
    - Genus Pygmephoridae
      - Pygmephoridae gen. sp. Batra,1965
    - Genus Pygmephorus
      - Pygmephorus spinosus Kramer, 1877 [Apis]
    - Genus Rhinopygmephorus
      - Rhinopygmephorus rhopalomelissae Kurosa, 2001
  - Family Scutacaridae Oudemans, 1916
    - Genus Imparipes
      - Imparipes (Apidacarus) apidophilus Mahunka, 1974
      - Imparipes (Apidacarus) paulyi Ebermann and Fain, 2002
      - Imparipes (Imparipes) apicola (Banks, 1914) (=Imparipes americanus) [Apis]
      - Imparipes (Imparipes) breganti Hall and Ebermann, 2004
      - Imparipes (Imparipes) eickworti Mahunka, 1969
      - Imparipes (Imparipes) floridensis Delfinado and Baker, 1976
      - Imparipes (Imparipes) haeseleri Ebermann and Hall, 2004
      - Imparipes (Imparipes) ithacensis Delfinado and Baker, 1976
      - Imparipes (Imparipes) mexicanus Delfinado and Baker, 1976
      - Imparipes (Imparipes) neotropicus Delfinado and Baker, 1976
      - Imparipes (Imparipes) rafalskii Dastych, 1978 [Bombus]
      - Imparipes (Imparipes) texanus Cockerell, 1910
      - Imparipes (Imparipes) vulgaris Delfinado and Baker, 1976
      - Imparipes dispar Rack, 1964
      - Imparipes parapicola Delfinado, Baker and Abbatiello, 1976
      - Imparipes sp. Ordway, 1964
    - Genus Nasutiscutacarus
      - Nasutiscutacarus ampliatus Beer and Cross, 1960
      - Nasutiscutacarus anthrenae Beer and Cross, 1960
    - Genus Parascutacarus
      - Parascutacarus indicus Baker and Delfinado, 1975 [Bombus]
    - Genus Scutacarus
      - Scutacarus (Scutacarus) eickworti Delfinado and Baker, 1976
      - Scutacarus acarorum (Goeze, 1780) [Apis Bombus]
      - Scutacarus deserticolus Mahunka, 1969 [Bombus]
      - Scutacarus mendax Karafiat, 1959 [Bombus]
      - Scutacarus occultatus Sevastianov, 1975 (=Scutacarus sinacarorum Dobrev and Mahunka, 1990) [Bombus]
  - Family Stigmaeidae Oudemans, 1931
    - Genus Storchia
      - Storchia sp. Malabanan & Corpuz-Raros, 1988 [Apis]
  - Family Tarsonemidae Kramer, 1877
    - Genus Acarapis
      - Acarapis dorsalis Morgenthaler, 1934 [Apis]
      - Acarapis externus Morgenthaler in Morison, 1931 [Apis]
      - Acarapis vagans Schneider, 1941 (nomen dubium) [Apis]
      - Acarapis woodi (Rennie, 1921) [Apis Bombus]
    - Genus Crossacarapis
      - Crossacarapis eickworti Ochoa and OConnor, 1996
    - Genus Dendroptus
      - Dendroptus sp. Sumangala et al., 2001 [Apis]
    - Genus Pseudacarapis
      - Pseudacarapis indoapis (Lindquist, 1968) [Apis]
      - Pseudacarapis trispicula Ochoa and Pettis, 2003 [Apis]
    - Genus Tarsonemus
      - Tarsonemus apis Rennie, 1921 [Apis]
      - Tarsonemus blakemorei Delfinado, 1978 [Apis]
      - Tarsonemus fusarii Cooreman, 1941 [Apis Bombus]
      - Tarsonemus minimax Vitzthum, 1926 [Apis]
      - Tarsonemus nr. aegualis Livshits, Mitrofanov and Sharonov, 1979 [Apis]
      - Tarsonemus platynopodae Magowski, 1986
      - Tarsonemus spp. Haragsim at al., 1978; De Jong et al., 1985;Malabanan&Corpuz-Raros, 1988; Kamali at al., 2001 [Apis]
      - Tarsonemus xylocopae Magowski, 1986
  - Family Tenuipalpidae Berlese, 1913
    - Genus Bryobia
      - Bryobia rubrioculus (Scheuten, 1857) (=Bryobia redicorzevi Reck) [Apis]
      - Bryobia spp. Kamali at al., 2001); Davis and McRory, 1987 [Apis]
    - Genus Cenopalpus
      - Cenopalpus sp. Kamali at al., 2001 [Apis]
  - Family Tetranychidae Donnadie, 1875
    - Genus Tetranychidae
      - Tetranychidae gen. spp. Chmielewski, 1991; Grobov, 1975 [Apis]
    - Genus Tetranychus
      - Tetranychus sp. Banaszak, 1980 [Apis]
      - Tetranychus urticae C. L. Koch, 1836 [Apis]
      - Tetranychus urticae C. L. Koch, 1836 sp. complex [Apis]
  - Family Trochometridiidae Mahunka, 1970
    - Genus Trochometridium
      - Trochometridium iranicum Hajiqanbar and Khaustov, 2009 [Apis]
      - Trochometridium tribulatum Cross, 1965
  - Family Trombiculidae Ewing, 1929
    - Genus Trombiculidae
      - Trombiculidae sp. Chmielewski, 1971 [Apis]
  - Family Trombidiidae Leach, 1815
    - Genus Allothrombium
      - Allothrombium fuliginosum (Hermann, 1804) [Apis]
    - Genus Trombidium
      - Trombidium holosericum (Linnaeus, 1758) [Apis]
  - Family Tydeidae Kramer, 1877
    - Genus Melissotydeus
      - Melissotydeus macrosolenus André, 1985
    - Genus Paralorryia
      - Paralorryia sp. Mosaddegh, 1997 [Apis]
    - Genus Proctotydaeus
    - Genus Pronematus
      - Pronematus sp. Malabanan and Corpuz-Raros, 1988 [Apis]
    - Genus Tydeidae
      - Tydeidae gen. sp. Chmielewski, 1991; de Jong, 1982 [Apis]
    - Genus Tydeus
      - Tydeus interruptus Thor, 1932 [Bombus]
      - Tydeus sp. Haragsim at al., 1978; Malabanan and Corpuz-Raros, 1988; Homann, 1933; Chmielewski, 1971; Kamali at al., 2001 [Apis Bombus]
      - Tydeus xylocopae Oudemans, 1926

PARASITIFORMES Reuter, 1909

Ixodida Leach, 1815
- - Family Ixodidae Dugès, 1834
    - Genus Dermacentor
      - Dermacentor variabilis (Say, 1821)
    - Genus Haemaphysalis
      - Haemaphysalis concinna C. L. Koch, 1844 [Apis]
    - Genus Hyalomma
      - Hyalomma sp. Grobov, 1978 [Apis]
    - Genus Ixodes
      - Ixodes sp. Grobov, 1975, 1978 [Apis]
Mesostigmata G. Canestrini, 1891
- Monogynaspida Camin & Gorirossi, 1955
  - Family 'Delfinadoidae'
    - Genus 'Delfinado
      - 'Delfinado sp.' Putatunda and Arbol, 2003
  - Family Ameroseiidae Evans, 1961
    - Genus Afrocypholaelaps
      - Afrocypholaelaps africana (Evans, 1963) [Apis]
      - Afrocypholaelaps analicullus Ho, Ma,Wang, Severinghaus, 2010 [Apis]
    - Genus Ameroseius
      - Ameroseius bembix ealensis Elsen, 1973
      - Ameroseius benoiti Elsen, 1973
      - Ameroseius plumigerus (Oudemans, 1930) [Apis]
      - Ameroseius plumosus (Oudemans, 1902) [Apis Bombus]
      - Ameroseius sp. Banaszak, 1980; Davydova & Bogatyrev, 1990 [Apis Bombus]
    - Genus Hattena
      - Hattena dalyi (Elsen, 1974)
      - Hattena ewae (Haitlinger, 1987)
      - Hattena tonganus (Mason, 1974)
    - Genus Neocypholaelaps
      - Neocypholaelaps ampullula (Berlese, 1910) [Apis]
      - Neocypholaelaps apicola Delfinado-Baker and Baker, 1983 [Apis]
      - Neocypholaelaps breviperitremata Elsen, 1972
      - Neocypholaelaps cocos Evans, 1963 [Apis]
      - Neocypholaelaps crocisae Elsen, 1972
      - Neocypholaelaps favus Ishikawa, 1968 [Apis]
      - Neocypholaelaps indica Evans, 1963 [Apis]
      - Neocypholaelaps leopoldi Elsen, 1972
      - Neocypholaelaps nova Elsen, 1972
      - Neocypholaelaps novaehollandiae Evans, 1963 [Apis]
      - Neocypholaelaps phooni Baker and Delfinado-Baker, 1985
      - Neocypholaelaps sp. Putatunda and Arbol, 2003 [Apis]
      - Neocypholaelaps stridulans (Evans, 1955) [Apis]
      - Neocypholaelaps varipilosa Elsen, 1972
      - Neocypholaelaps xylocopae Elsen, 1972
    - Genus Sertitympanum
      - Sertitympanum aegyptiacus Naser and Abow-Awad, 1986 [Apis]
  - Family Ascidae Voigts & Oudemans, 1905
    - Genus Antennoseius
      - Antennoseius spp. Kamali at al., 2001;Mosaddegh, 1997 [Apis]
    - Genus Arctoseius
      - Arctoseius cetratus (Sellnick, 1940) [Apis]
  - Family Blattisociidae Garman, 1948
    - Genus Blattisocius
      - Blattisocius apis Basha and Yousef, 2000 [Apis]
      - Blattisocius apisassociae Chinniah and Mohanasundaram, 1995 [Apis]
      - Blattisocius dentriticus (Berlese, 1918) [Apis]
      - Blattisocius keegani (Fox, 1947) [Apis]
      - Blattisocius mali (Oudemans, 1929) [Apis]
      - Blattisocius tarsalis (Berlese, 1918) [Apis]
  - Family Ascidae Voigts & Oudemans, 1905
    - Genus Gamasellodes
      - Gamasellodes bicolor (Berlese, 1918) [Apis]
  - Family Blattisociidae Garman, 1948
    - Genus Lasioseius
      - Lasioseius berlesei (Oudemans, 1938) [Apis]
      - Lasioseius floridensis Berlese group [Apis]
      - Lasioseius furcisetus Athias-Henriot, 1959 [Apis]
      - Lasioseius krantzi Chant, 1963 [Apis]
      - Lasioseius penicilliger (Berlese, 1916) [Apis Bombus]
      - Lasioseius spp. De Jong et al., 1985; Malabanan and Corpuz-Raros, 1988 [Apis]
  - Family Melicharidae Hirschmann, 1962
    - Genus Melichares
      - Melichares agilis Hering, 1838 [Apis]
    - Genus Proctolaelaps
      - Proctolaelaps bickleyi (Bram, 1956) [Apis]
      - Proctolaelaps bombophilus (Postner in Westerboer, 1963) [Bombus]
      - Proctolaelaps longanalis (Westerboer, 1963) [Bombus]
      - Proctolaelaps longisetosus (Postner in Westerboer, 1963) (=Proctolaelaps novineus El-Banhawy and Nasr, 1986) [Bombus]
      - Proctolaelaps ornatus (Postner in Westerboer, 1963) [Bombus]
      - Proctolaelaps pygmaeus (Müller, 1859) [Apis Bombus]
      - Proctolaelaps scolyti Evans, 1958 (=Proctolaelaps ulmi Hirsch) [Apis]
      - Proctolaelaps sibiriensis (Davydova, 1988) [Bombus]
    - Genus Spadiseius
      - Spadiseius spathiphyllae Lindquist and Moraza 2008
  - Family Dermanyssidae Kolenati, 1859
    - Genus Dermanyssus
      - Dermanyssus gallinae (DeGeer, 1778) [Apis]
  - Family Dinychidae Berlese, 1916
    - Genus Dinychus
      - Dinychus sp. Davydova et al., 1987 [Bombus]
  - Family Eviphididae Berlese, 1913
    - Genus Alliphis
      - Alliphis halleri (G. Canestrini and R. Canestrini, 1881) [Apis]
      - Alliphis siculus (Oudemans, 1905) [Apis]
    - Genus Eviphis
      - Eviphis spp. De Jong et al., 1985; Grobov, 1974; Homann, 1933 [Apis]
  - Family Haemogamasidae Oudemans, 1926
    - Genus Eulaelaps
      - Eulaelaps stabularis (Koch, 1839) [Apis]
    - Genus Haemogamasus
      - Haemogamasus mandschuricus Vitzthum, 1930 [Apis]
      - Haemogamasus nidi Michael, 1892 [Apis]
      - Haemogamasus nidiformis Bregetova, 1956 [Bombus]
      - Haemogamasus pontiger (Berlese, 1903) [Apis]
    - Genus Hirstionyssus
      - Hirstionyssus sp. Kamali at al., 2001; Mosaddegh, 1997 [Apis]
  - Family Halolaelapidae Karg, 1965
    - Genus Halolaelaps
      - Halolaelaps saproincisus (Hirschmann and Gotz, 1968) [Apis]
      - Halolaelaps sp. Mosaddegh, 1997 [Apis]
  - Family Laelapidae Berlese, 1892
    - Genus Androlaelaps
      - Androlaelaps bayoumi Basha and Yousef, 2000 [Apis]
      - Androlaelaps casalis (Berlese, 1887) [Apis]
      - Androlaelaps fahrenholzi (Berlese, 1911) (=Androlaelaps glasgowi (Ewing, 1925)) [Apis]
      - Androlaelaps myrmecophila (Evans and Till, 1966) [Bombus]
    - Genus Bisternalis
      - Bisternalis camargoi Baker, Flechtmann and Delfinado-Baker 1984
      - Bisternalis formosus Baker, Flechtmann and Delfinado-Baker 1984
      - Bisternalis mexicanus Baker, Delfinado-Baker and Reyes Ordaz, 1983
      - Bisternalis rettenmeyeri Hunter, 1963
      - Bisternalis trigonarum Baker, Flechtmann and Delfinado-Baker 1984
    - Genus Cosmolaelaps
      - Cosmolaelaps vacuus (Michael, 1891) [Bombus]
    - Genus Dinogamasus
      - Dinogamasus acutus LeVeque 1930
      - Dinogamasus affinis (Berlese 1918)
      - Dinogamasus albulus Lundqvist, 1998
      - Dinogamasus alfkeni (Oudemans 1902)
      - Dinogamasus amaniensis (Vitzthum 1919)
      - Dinogamasus assimiensis Lundqvist, 1998
      - Dinogamasus bakeri LeVeque 1931
      - Dinogamasus bequaerti LeVeque 1930
      - Dinogamasus braunsi (Vitzthum 1914)
      - Dinogamasus brevihirtus LeVeque 1930
      - Dinogamasus brevipes LeVeque 1931
      - Dinogamasus collarti (Oudemans 1929)
      - Dinogamasus concinnus LeVeque 1931
      - Dinogamasus crassipes Kramer 1898
      - Dinogamasus heteraspis LeVeque 1930
      - Dinogamasus inflatus LeVeque 1930
      - Dinogamasus jacobsoni (Berlese 1910)
      - Dinogamasus kerrianus LeVeque 1931
      - Dinogamasus kordofaniensis Lundqvist, 1998
      - Dinogamasus levequae Lundqvist, 1998
      - Dinogamasus macgregori LeVeque 1931
      - Dinogamasus maxima (Vitzthum 1919)
      - Dinogamasus minor Lundqvist, 1998
      - Dinogamasus northolmensis Loots 1980
      - Dinogamasus occidentalis Lundqvist, 1998
      - Dinogamasus octoconus LeVeque 1931
      - Dinogamasus oudemansi LeVeque 1930
      - Dinogamasus parvus LeVeque 1930
      - Dinogamasus perkinsi (Oudemans, 1901)
      - Dinogamasus philippinensis LeVeque 1930
      - Dinogamasus piperi LeVeque 1930
      - Dinogamasus productus LeVeque 1930
      - Dinogamasus ramaleyi LeVeque 1931
      - Dinogamasus similis LeVeque 1931
      - Dinogamasus sternisetosa (Vitzthum 1930)
      - Dinogamasus tonkinensis Lundqvist, 1998
      - Dinogamasus tortivus Lundqvist, 1998
      - Dinogamasus trihirtus LeVeque 1931
      - Dinogamasus villosior (Berlese 1918)
    - Genus Eumellitiphis
      - Eumellitiphis inouei Delfinado-Baker and Baker, 1988
      - Eumellitiphis mellitus Türk, 1948
      - Eumellitiphis philippinensis Delfinado-Baker and Baker, 1988
    - Genus Euvarroa
      - Euvarroa sinhai Delfinado and Baker, 1974 [Apis]
      - Euvarroa wongsirii Lekprayoon and Tangkanasing, 1991 [Apis]
    - Genus Hunteria
      - Hunteria brasiliensis Delfinado-Baker, Baker and Flechtmann, 1984
    - Genus Hypoaspis
      - Hypoaspis alphabetica Berlese, 1904
      - Hypoaspis asperatus (Berlese, 1905) [Apis]
      - Hypoaspis favosa Türk, 1948
      - Hypoaspis heselhausi Oudemans, 1912 [Bombus]
      - Hypoaspis hoffmannae Smiley, Baker and Delfinado-Baker, 1996
      - Hypoaspis lubrica Voigts and Oudemans, 1904 [Apis Bombus]
      - Hypoaspis meliponarum Vitzthum, 1930
      - Hypoaspis nr. lubrica Voigts and Oudemans, 1904 [Apis]
    - Genus Laelaps
      - Laelaps algericus Hirst, 1925 [Apis]
    - Genus Laelaspoides
      - Laelaspoides ordwayae Eickwort, 1966
    - Genus Meliponaspis
      - Meliponaspis debilipes Vitzthum, 1930
    - Genus Melittiphis
      - Melittiphis alvearius (Berlese, 1895) [Apis]
    - Genus Melittiphisoides
      - Melittiphisoides apiarium Delfinado-Baker, Baker and Flechtmann, 1984
    - Genus Neohypoaspis
      - Neohypoaspis ampliseta Delfinado-Baker, Baker and Roubik, 1983
    - Genus Pneumolaelaps
      - Pneumolaelaps 'bombicolens (Canestrini)' [Apis]
      - Pneumolaelaps 'brevista Evans and Till' [Apis]
      - Pneumolaelaps aequalipilus Hunter, 1966 [Bombus]
      - Pneumolaelaps bombicolens (Canestrini, 1884) [Apis Bombus]
      - Pneumolaelaps breviseta (Evans and Till, 1966) [Bombus]
      - Pneumolaelaps cf. patae Hunter and Husband, 1973
      - Pneumolaelaps colomboi (Evans and Till 1966) [Apis Bombus]
      - Pneumolaelaps connieae Hunter and Husband, 1973 [Bombus]
      - Pneumolaelaps costai Hunter and Husband, 1973 [Bombus]
      - Pneumolaelaps fuscicolens (Oudemans, 1903) [Bombus]
      - Pneumolaelaps groenlandicus (Trägårdth, 1904) [Bombus]
      - Pneumolaelaps hyatti (Evans and Till, 1966) (=Qinghailaelaps gui Bai, 1992) [Apis Bombus]
      - Pneumolaelaps longanalis Hunter and Husband, 1973 [Apis Bombus]
      - Pneumolaelaps longipilus Hunter, 1966 [Bombus]
      - Pneumolaelaps machadoi Elsen, 1973
      - Pneumolaelaps marginepilosus (Sellnick, 1938) [Apis Bombus]
      - Pneumolaelaps minutissima (Evans and Till, 1966) [Bombus]
      - Pneumolaelaps mistipilus Hunter, 1966 [Bombus]
      - Pneumolaelaps nr. richardsi (Hunter and Husband) [Apis]
      - Pneumolaelaps patae Hunter and Husband, 1973 [Bombus]
      - Pneumolaelaps qinghaiensis (Li, Yang and Wang, 1998) [Bombus]
      - Pneumolaelaps richardsi Hunter and Husband, 1973 [Bombus]
      - Pneumolaelaps sinhai Hunter and Husband, 1973 [Apis Bombus]
      - Pneumolaelaps sp. Garrido, 1995 [Bombus]
    - Genus Raymentia
      - Raymentia anomala Womersley, 1956
      - Raymentia eickwortiana Walter and Beard, 2002
      - Raymentia walkeriana Walter and Beard, 2002
    - Genus Stevelus
      - Stevelus amiculus Hunter, 1963
    - Genus Stigmatolaelaps
      - Stigmatolaelaps greeni (Oudemans, 1902)
      - Stigmatolaelaps hunteri Krantz, 1998
      - Stigmatolaelaps sumatrensis Krantz, 1998
    - Genus Stratiolaelaps
      - Stratiolaelaps miles (Berlese, 1892) [Apis]
    - Genus Tropilaelaps
      - Tropilaelaps clareae Delfinado and Baker, 1962 [Apis]
      - Tropilaelaps koenigerum Delfinado-Baker and Baker, 1982 [Apis]
      - Tropilaelaps mercedesae Anderson and Morgan, 2007 [Apis]
      - Tropilaelaps thaii Anderson and Morgan, 2007 [Apis]
    - Genus Urozercon
      - Urozercon angustatus Silvestri, 1911, stat. n.
      - Urozercon melittophilus Silvestri, 1911
      - Urozercon sp. Klimov (unpubl.)
    - Genus Varroa
      - Varroa destructor Anderson and Trueman, 2000 [Apis]
      - Varroa jacobosoni Oudemans, 1904 [Apis]
      - Varroa rindereri de Guzman and Delfinado-Baker, 1996 [Apis]
      - Varroa underwoodi Delﬁnado-Baker and Aggarwal, 1987 [Apis]
    - Genus Xylocolaelaps
      - Xylocolaelaps burgetti Royce and Krantz, 2003
    - Genus Zontia
      - Zontia meliponensis Türk, 1948
  - Family Macrochelidae Vitzthum, 1930
    - Genus Glyptholaspis
      - Glyptholaspis americana (Berlese, 1888) [Apis]
    - Glyptholaspis asperrima (Berlese, 1905) [Apis]
  - Glyptholaspis pontina Filipponi and Pegazzano, 1960 [Apis]
- Genus Holostaspella
  - Holostaspella pulchella Mašán, 2003 [Apis]
- Genus Macrocheles
  - Macrocheles caucasicus Bregetova and Koroleva, 1960 [Apis]
  - Macrocheles decoloratus (C.L. Koch, 1839) [Apis]
  - Macrocheles dispar (Berlese, 1910) [Apis]
  - Macrocheles glaber (Muller, 1860) [Apis]
  - Macrocheles hallidayi Walter and Krantz, 1986 [Apis]
  - Macrocheles kraepelini (Berlese, 1905) [Apis]
  - Macrocheles limue Samšinăk, 1962 [Apis]
  - Macrocheles mammifer Berlese, 1918 [Apis]
  - Macrocheles martius (Hull, 1925) [Apis]
  - Macrocheles muscaedomesticae (Scopoli, 1772) [Apis]
  - Macrocheles nataliae Bregetova and Koroleva, 1960 [Apis]
  - Macrocheles nidus Hartini, Kahono and Takaku, 2013 [Apis]
- Macrocheles penicilliger (Berlese, 1904) [Bombus]
Macrocheles praedafimetorum Richards and Richards, 1977 [Apis Bombus]

Macrocheles rotundiscutis Bregetova and Koroleva, 1960 (=Macrocheles bombophilus Goetz, in Hirschmann, 1970) [Bombus]

Genus Neopodocinum

Neopodocinum halimunense Hartini and Takaku, 2003 [Apis]

Genus Trigonholaspis

Family Macronyssidae Oudemans, 1936

Genus Macronyssidae
- Macronyssidae sp. Kamali at al., 2001; Mosaddegh, 1997 [Apis]

Family Microgyniidae Trägårdh, 1942

Genus Microgyniidae
- Microgyniidae sp. Grobov, 1978 [Apis]

Family Ologamasidae Ryke, 1962

Genus Cyrtolaelaps
- Cyrtolaelaps mucronatus (G. and R. Canestrini 1881) [Apis]
- Cyrtolaelaps sp. Grobov, 1978 [Apis]
Genus Euryparasitus
- Euryparasitus medius Zuevsky, 1971 [Bombus]
Genus Gamasiphis
- Gamasiphis sp. Grobov, 1978 [Apis]

Family Oplitidae Johnston, 1968

Genus Oplitis
- Oplitis (Uroplitana) meliponarum Türk, 1948
- Oplitis minutissima (Berlese, 1903) [Apis]

Family Pachylaelapidae Berlese, 1913

Genus Meliponapachys
- Meliponapachys pallidus Türk, 1948 (=Tentative family assignment)

Family Parasitidae Oudemans, 1901

Genus Eugamasus
- Eugamasus sp. Mosaddegh, 1997 [Apis]
Genus Holoparasitus
- Holoparasitus sp. Grobov, 1978
Genus Parasitellus
- Parasitellus arcticus (Karg, 1985) [Bombus]
- Parasitellus crinitus (Oudemans, 1903) [Bombus]
- Parasitellus fucorum (De Geer, 1778) [Apis Bombus]
- Parasitellus hobbsi (Richards, 1976) [Apis Bombus]
- Parasitellus ignotus (Vitzthum, 1930) [Bombus]
- Parasitellus inquilinobombus (Richards, 1976) [Bombus]
- Parasitellus jamalensis Davydova, 1988 [Bombus]
- Parasitellus netskyi (Davydova and Bogdanov, 1976) [Bombus]
- Parasitellus nr. crinitus (Oudemans, 1903) [Apis]
- Parasitellus nr. perthecatus (Richards, 1976) [Apis]
- Parasitellus nr. talparum (Oudemans, 1913) [Apis]
- Parasitellus papei Karg, 1985 [Bombus]
- Parasitellus perthecatus (Richards, 1976) [Bombus]
- Parasitellus sp1 Crozier, 1989 [Apis Bombus]
- Parasitellus sp2 Klimov (unpubl.) [Bombus]
- Parasitellus talparum (Oudemans, 1913) (=Parasitus anglicus Vitzthum 1930; Parasitus bombophilus Vitzthum 1930; Parasitus numismaticus Vitzthum 1930; Parasitus favus Richards, 1976; Parasitus traegardhi Micherdzinski, 1969; Parasitus crinitosimilis Vitzthum, 1930) [Bombus]
Genus Parasitus
- Parasitus beta Oudemans and Voigts, 1904 [Apis Bombus]
- Parasitus coleoptratorum (Linnaeus, 1758) [Apis Bombus]
- Parasitus consanguineus (Oudemans and Voigts, 1904) [Bombus]
- Parasitus fimetorum Berlese, 1904 [Apis Bombus]
- Parasitus loricatus (Wankel, 1861) [Bombus]
- Parasitus sp. Haragsim at al., 1978; De Jong et al., 1985 [Apis]
Genus Pergamasus
- Pergamasus crassipes (Linnaeus, 1758) [Apis]
- Pergamasus diversus Halbert, 1915 [Apis]
Genus Poecilochirus
- Poecilochirus necrophori Vitzthum, 1930 [Apis]
- Poecilochirus subterraneus (Müller, 1860) [Apis]
Genus Vulgarogamasus
- Vulgarogamasus aff. kraepelini (Berlese, 1904) [Apis]
- Vulgarogamasus cornutosimilis (Schweizer, 1949) [Bombus]
- Vulgarogamasus pollinerus El-Banhawy and Nasr, 1984 [Apis Bombus]

Family Phytoseiidae Berlese, 1916

Genus Amblydromella
- Amblydromella kettanehi (Dosse, 1967) [Apis]
Genus Amblyseius
- Amblyseius (Neoseiulus) marginatus (Wainstein, 1961) [Apis]
- Amblyseius spp. Kumar, 1996; Kamali at al., 2001; Mosaddegh, 1997 [Apis]
Genus Fundiseius
- Fundiseius timagami (Chant and Hansell, 1971) [Apis]
Genus Neoseiulus
- Neoseiulus barkeri Hughes, 1948 [Apis]
- Neoseiulus cucumeris (Oudemans, 1930) [Bombus]
- Neoseiulus nr. 'swarti Zack' [Apis]
Genus Proprioseiopsis
- Proprioseiopsis okanagensis (Chant, 1957) [Apis]
Genus Typhlodromus
- Typhlodromus bakeri (Garman, 1948) [Apis]
- Typhlodromus diumbokus Schicha and Corpuz-Raros, 1992 [Apis]

Family Polyaspididae Berlese, 1913

Genus Polyaspis
- Polyaspis sp. Sumangala et al., 2001 [Apis]

Family Rhodacaridae Oudemans, 1902

Genus Rhodacarus
- Rhodacarus sp. Kamali at al., 2001; Mosaddegh, 1997 [Apis]

Family Trachytidae Trägårdh, 1938

Genus Trachytes
- Trachytes aegrota (C.L.Koch, 1841)
Genus Uroseius
- Uroseius acuminatus (C. L. Koch, 1842) [Apis]
- Uroseius jabae Berlese, 1910 [Apis]

Uroseius hunzikeri Schweizer, 1922 [Bombus]

Family Trematuridae Berlese, 1917

Genus Nenteria
- Nenteria breviunguiculata (Willmann, 1949)
- Nenteria eulaelaptis (Vitzthum, 1930)
- Nenteria postneri Hirschmann and Zirngiebl-Nicol 1969 [Bombus]
Nenteria stlylifera (Berlese, 1904)

Genus Trematura

Trematura nr bassusi Wiśniewski and Hirschmann, 1987

Genus Trichouropoda

Trichouropoda parasítica (Choudhuri and Mukherjee, 1964) [Apis]

Trichouropoda stercoraria Hirschmann and Hiramatsu, 1990 [Apis]

Genus Trichosociata

Trichosociata nr peritrematalis (Hirschmann, 1972)

Genus Pseuduropoda

Pseuduropoda sp. Klimov, unpubl.

Family Uroactiniidae Hirschmann & Zirngiebl-Nicol, 1964

Genus Uroactinia
- Uroactinia centroamericana (Stoll 1893)

Family Urodinychidae Berlese, 1917

Genus Urodiaspis
- Urodiaspis tecta (Kramer, 1876)
Genus Uroobovella
- Uroobovella (Fuscouropoda) sp. Çankaya&Kaftanoglu, 2006 [Bombus]
- Uroobovella marginata (C.L. Koch, 1839) [Apis Bombus]

Uroobovella sp. Macfarlane, 2005 [Bombus]

Family Uropodidae Kramer, 1881

Genus Discourella
- Discourella cordieri (Berlese, 1916) [Bombus]
Genus Uropoda
- Uropoda sp. Piletskaya and Zaloznaya, 2004 [Apis]
- Uropoda sp. Gordh & Barrows, 1976
- Uropoda orbicularis (Müller, 1776)
Uropoda (Uropoda) minima Kramer, 1882

Genus Uropodidae

Uropodidae sp. Delfinado-Baker et al., 1991, Chmielewski, 1971; Davydova et al., 1987 [Apis Bombus]

Family Veigaiidae Oudemans, 1939

Genus Veigaia
- Veigaia kochi (Trägårdh, 1901) [Bombus]

Veigaia nemorensis (C. H. Koch, 1838) [Apis]

Family Zerconidae G. Canestrini, 1891

Genus Zerconidae
- Zerconidae sp. Grobov, 1978 [Apis]

Family Sejidae Berlese, 1885

Genus Epicroseius
- Epicroseius nr. tanganicus (Hirschmann and Kaczmarek 1991) [Apis]

Family Diplogyniidae Trägårdh, 1941

Genus Celaenosthanus
- Celaenosthanus trigonophilus Vitzthum, 1930
- Celaenosthanus sp1 Kethley, 1977

Celaenosthanus sp2 Kethley, 1977

Family Physalozerconidae

Genus Physalozercon
- Physalozercon raffray (Wasmann, 1902) [Bombus]

Family Triplogyniidae Funk, 1977

Genus Funkotriplogynium
- Funkotriplogynium irapora (Flechtman, 1983)