Conventions:
1) Each character is separated from the next by double paragraph marks.
2) Each character is separated from the respective character states by a paragraph mark.
3) Each character state is separated from the next by a paragraph marks.
4) Figures are linked to character states using the following expression: Fig-ch0-0.jpg, where "ch0-0.jpg" is the name of the figure file located in the folder named "Figs".
5) Maps are linked to character states using the following expression: Map-DiadasiopusV.gif?67,91,110,135. Where "Map-DiadasiopusV.gif " is the name of the figure file located in the folder named "Maps". 67,91,110,135 is the leftmost (X), topmost (Y) position, width, and height in pixels of the frame representing the corresponding area on the character map.
An example of the format is below (numbering is optional. Any html tags can be used to format text; for example, the expression <u>or</u> underlines the world "or")
Characters and Character States from the Key to Families of Bee Mites
0. Coxae
0) fused to ventral body forming coxal fields Fig-ch0-0.jpg Map-DiadasiopusV.gif?67,91,110,135
1) free; coxae I separated, not overlapping or covering tritosternal base Fig-ch0-1.jpg Map-MellitiphisV.gif?113,85,86,92
2) free; coxae I nearly touching each other, partially overlapping or covering tritosternal base Fig-ch0-2.jpg
1. Tritosternum
0) absent Fig-ch1-0.jpg
1) present Fig-ch1-1.jpg
2. Posterior ventral region
0) without suckers, or with 1 pair in males Fig-ch2-0.jpg Map-MedeusV.gif?53,164,113,116
1) with suckers and conoids organized in attachment organ Fig-ch2-1.jpg Map-DiadasiopusV.htm?67,152,107,117
3. Gnathosoma
0) not reduced, functional; if reduced then male with genital capsule present Fig-ch3-0.jpg Map-MedeusV.gif?75,16,74,77
1) variously reduced, non-functional; immatures with no external genitalia Fig-ch3-1.jpg Map-DiadasiopusV.htm?82,43,74,79
4. Terminal solenidion of palp
0) shorter than free palpomeres Fig-ch4-0.jpg Map-MedeusV.gif?92,21,40,43
1) longer than free palpomeres Fig-ch4-1.jpg Map-DiadasiopusV.htm?87,41,64,60
2) absent Fig-ch4-2.jpg Map-DiadasiopusV.htm?87,41,64,60
3) inaplicable because palps absent Fig-ch4-3.jpg Map-DiadasiopusV.htm?103,50,33,33
5. Attachment organ
0) well-developed, exceeding 1/5 of body width, with at least 2 pairs of suckers (ad1+2 and ad3) and 2 pairs conoids (ps1 and ps2) Fig-ch5-0.jpg Map-DiadasiopusV.htm?70,153,100,114
1) vestigial, less than 1/5 of body width, with only 1-2 distinct suckers Fig-ch5-1.jpg Map-DiadasiopusV.htm?82,194,85,73
2) absent (1 pair of suckers may be present in some male Astigmata) Fig-ch2-0.jpg Map-MedeusV.gif?52,169,122,110
6. Solenidion omega1
0) situated on tibia I close to solenidion phi Fig-ch6-0.jpg Map-DiadasiopusD.htm?70,25,32,28
1) situated on tarsus I Fig-ch6-1.jpg Map-DiadasiopusD.htm?67,7,35,34
2) absent Fig-ch6-2.jpg
3) unknown, because all leg I podomeres fused and leg I with one solenidion Fig-ch6-3.jpg Map-DiadasiopusV.htm?70,15,39,70
7. Legs IV in repose
0) flexed at trochanter-femur articulation and directed anteriorly Fig-ch7-0.jpg Map-DiadasiopusV.htm?46,86,177,185
1) directed posteriorly or laterally Fig-ch7-1.jpg Map-DiadasiopusV.htm?41,119,164,149
2) absent
8. Medial articulation of tarsi IV
0) present Fig-ch8-0.jpg Map-DiadasiopusV.htm?149,211,48,48
1) absent Fig-ch8-1.jpg Map-DiadasiopusV.htm?149,211,48,48
2) legs IV absent
9. Setae ve
0) present Fig-ch9-0.jpg Map-DiadasiopusD.htm?85,47,70,73
1) absent Fig-ch9-1.jpg Map-DiadasiopusD.htm?85,47,70,73
10. Setae vi
0) present Fig-ch10-0.jpg Map-DiadasiopusD.htm?74,40,88,80
1) absent Fig-ch10-1.jpg Map-DiadasiopusD.htm?102,47,33,28
11. Setae vi situated
0) near anterior edge of propodosoma Fig-ch11-0.jpg Map-MedeusD.gif?69,6,97,99
1) about half-way between anterior edge of propodosoma and external scapular setae se Fig-ch11-1.jpg Map-MedeusD.gif?69,6,97,99
2) at level of scapular setae or posteriorly
3) absent/non-applicable
12. Genu and tibia III and IV (if leg IV present)
0) nearly equal or genu longer or less than 4 times shorter than corresponding tibia Fig-ch12-0.jpg Map-DiadasiopusV.htm?159,188,40,41
1) genu more than 4 times shorter than corresponding tibia Fig-ch12-1.jpg Map-DiadasiopusV.htm?159,188,40,41
2) unknown because leg podomeres fused Fig-ch12-2.jpg Map-DiadasiopusV.htm?159,188,40,41
13. Pretarsal empodial claw III
0) present Fig-ch13-0.jpg Map-DiadasiopusV.htm?159,188,42,42
1) absent Fig-ch13-1.jpg Map-DiadasiopusV.htm?159,188,42,42
14. Pretarsal empodial claw IV
0) present Fig-ch14-0.jpg Map-DiadasiopusV.htm?153,215,40,40
1) absent Fig-ch14-1.jpg Map-DiadasiopusV.htm?153,215,40,40
15. Leg IV
0) nearly equal in length and similar to leg III Fig-ch15-0.jpg Map-DiadasiopusV.htm?35,151,124,112
1) distinctly shorter than leg III and with longer tarsal setae; some podomeres may be fused Fig-ch15-1.jpg Map-DiadasiopusV.htm?35,151,124,112
2) absent
16. Solenidion omega3 on tarsus I placed
0) apically, subapically Fig-ch16-0.jpg Map-DiadasiopusD.htm?66,6,40,37
1) in basal half of tarsus I, close to solenidion omega1 Fig-ch16-1.jpg Map-DiadasiopusD.htm?66,6,40,37
2) unknown, because all podomeres fused and there is single solenidion on legs Fig-ch6-3.jpg Map-DiadasiopusV.htm?70,15,39,70
17. Anterior coxal apodemes III inner ends
0) fused to posterior apodemes III (coxal fields III enclosed but separated from each other) Fig-ch17-0.jpg Map-DiadasiopusV.htm?77,147,88,109
1) fused to median apodeme Fig-ch17-1.jpg Map-DiadasiopusV.htm?77,147,88,109
2) ending freely Fig-ch17-2.jpg Map-DiadasiopusV.htm?77,147,88,109
3) absent Fig-ch17-3.jpg
4) not applicable, because coxae free Fig-ch0-1.jpg
18. Propodosomal ocelli or pigmented spots
0) absent Fig-ch18-0.jpg Map-DiadasiopusD.htm?75,51,87,69
1) present Fig-ch18-1.jpg Map-DiadasiopusD.htm?75,51,87,69
19. Propodosomal shield
0) covers only part of propodosoma, distinctly separated from hysterosomal shield (if present) Fig-ch19-0.jpg Map-DiadasiopusD.htm?40,56,158,223
1) covers all or almost all propodosoma, not separated from hysterosomal shield (if present); if anterior projection present then without striate margin Fig-ch19-1.jpg Map-DiadasiopusD.htm?39,52,161,149
2) completely covers propodosoma; forming anterior projection <u>with</u> striate margin Fig-ch19-2.jpg
3) absent/or body covered by entire dorsal shield
20. Setae scx
0) filiform, sometimes pilose, pectinate, branched, spiniform or bifurcated Fig-ch20-0.jpg Map-DiadasiopusD.htm?74,71,39,35
1) vestigial, rod-like Fig-ch20-1.jpg Map-DiadasiopusD.htm?74,71,39,35
2) absent
21. Empodial claw/pulvillus II
0) claw-like or variously modified but not spirally twisted Fig-ch21-0.jpg Map-DiadasiopusD.htm?19,51,27,24
1) spirally twisted Fig-ch21-1.jpg Map-DiadasiopusD.htm?19,51,27,24
2) absent
22. Empodial claw II arising
0) directly from tarsal apices Fig-ch22-0.jpg Map-DiadasiopusD.htm?13,38,38,38
1) from membranous ambulacra Fig-ch22-1.jpg Map-DiadasiopusD.htm?13,38,38,38
2) absent Fig-ch22-2.jpg Map-DiadasiopusD.htm?9,40,33,31
23. Pretarsi II with
0) 1 empodial claw Fig-ch23-0.jpg
1) 1 empodium <u>and</u> 2 pretarsal claws Fig-ch23-1.jpg
24. Lateral opisthosomal glands
0) absent Fig-ch24-0.jpg
1) present Fig-ch24-1.jpg Map-DiadasiopusD.htm?40,119,158,158
25. Chelicerae
0) chelate-dentate, with few well-sclerotized teeth Fig-ch25-0.jpg Map-MedeusD.gif?95,3,48,48
1) attenuate, fixed digit and its teeth membranous Fig-ch25-1.jpg Map-MedeusD.gif?95,3,48,48
2) with movable digit styletiform or hooklike Fig-ch25-2.jpg
3) absent
26. Chelicerae
0) separate
1) fused basally forming stylophore
2) fused to each other and with subcapitulum forming stylophore capsule Fig-ch26-2.jpg
3) absent
27. Palp
0) not reflexed outward Fig-ch27-0.jpg Map-MedeusV.gif?90,21,42,41
1) reflexed outward Fig-ch27-1.jpg Map-MedeusV.gif?90,21,42,41
2) absent
28. Eupathidial and normal setae on palp
0) eupathidial setae, if present, shorter than free palpomeres, normal setae present Fig-ch28-0.jpg Map-MedeusV.gif?92,21,40,43
1) at least 1 eupathidial seta present, usually longer than free palpomeres, normal setae absent Fig-ch28-1.jpg Map-MedeusV.gif?73,15,72,80
2) all setae absent, palp reduced or absent
29. Genital papillae
0) situated under genital valves; cylindrical, conical, or bell-shaped Fig-ch29-0.jpg Map-MedeusV.gif?86,140,48,55
1) disassociated from genital opening; large ring-like (sometimes irregularly shaped) areas on external cuticle Fig-ch29-1.jpg Map-MedeusV.gif?54,93,117,112
2) absent
30. Pretarsal condylophores (not to be confused with vertical sclerites of Mesostigmata)
0) short, thick, less than 4 times longer than width Fig-ch30-0.jpg Map-MedeusV.gif?165,48,38,35
1) long, thin, more than 4 times longer than width Fig-ch30-1.jpg Map-MedeusV.gif?165,48,38,35
2) absent
31. Condylophores (not to be confused with vertical sclerites of Mesostigmata)
0) free Fig-ch31-0.jpg Map-MedeusV.gif?165,48,38,35
1) fused basally forming Y-shaped structure Fig-ch31-1.jpg Map-MedeusV.gif?165,48,38,35
2) fused basally forming assymetrical structure with sucker in basal part in male Fig-ch31-2.jpg Map-MedeusV.gif?165,48,38,35
3) absent
32. Male tarsus IV
0) without sucker-like setae (some vental suckers of non-setal origin may be on pretarsus) Fig-ch32-0.jpg Map-MedeusD.gif?2,209,41,37
1) with 2, usually dorsal sucker-like setae Fig-ch32-1.jpg Map-MedeusD.gif?2,209,41,37
2) non applicable because tarsus IV absent
33. Male para-anal suckers
0) absent Fig-ch33-0.jpg Map-MedeusV.gif?47,141,134,139
1) present Fig-ch33-1.jpg Map-MedeusV.gif?47,136,131,135
34. Tectal seta tc'' of tarsus II (=e in Astigmata)
0) filiform, or lanceolate distally Fig-ch34-0.jpg Map-MedeusD.gif?178,40,39,39
1) spiniform Fig-ch34-1.jpg Map-MedeusD.gif?178,40,39,39
2) filiform, with saucer-like tip Fig-ch34-2.jpg Map-DiadasiopusD.gif?178,31,54,54
3) absent or inapplicable Map-MedeusD.gif?20,46,33,33
35. Tectal setae tc'' (=e) and tc' (=f) of tarsus II
0) tc'' (=e) shorter than tc' (=f) Fig-ch35-0.jpg Map-MedeusD.gif?20,46,33,33
1) approximatelty equal, or tc'' (e) longer Fig-ch35-1.jpg Map-MedeusD.gif?20,46,33,33
2) absent/non-applicable
36. Unguinal setae (u' or u'') and proral setae (p' or p'') of tarsus II
0) unguinal setae shorter than proral setae Fig-ch36-0.jpg Map-MedeusV.gif?27,41,29,28
1) unguinal setae longer and sometimes stouter than proral setae, or approximately equal Fig-ch36-1.jpg Map-MedeusV.gif?27,41,29,28
2) inaplicable, or one pair or all setae absent
37. Proral setae (p' or p'') II
0) filiform, sometimes with lanceolate tip Fig-ch37-0.jpg Map-MedeusV.gif?27,41,29,28
1) spinform Fig-ch37-1.jpg Map-MedeusV.gif?27,41,29,28
2) enlarged and claw-like Fig-ch37-2.jpg Map-MedeusV.gif?27,41,29,28
3) spatuliform Fig-ch37-3.jpg Map-MedeusV.gif?27,41,29,28
4) absent/non-applicable
38. Idiosomal shape
0) distinctly longer than wide Map-MedeusD.gif?34,48,171,236
1) rounded, length similar to width Fig-ch38-1.jpg Map-MedeusD.gif?34,48,171,236
39. Dorsal setae
0) simple or variously modified, but not pectinate or slightly pectinate Map-MedeusD.gif?34,48,171,236
1) at least some setae heavily pectinate or plumose Map-MedeusD.gif?34,48,171,236
2) absent
40. Supracoxal gland opening
0) smaller than distance between setae vi (if present) Map-MedeusD.gif?86,12,81,89
1) larger than distance between setae vi, placed on distinct oval sclerite above legs I Fig-ch40-1.jpg Map-MedeusD.gif?86,12,81,89
2) absent/non-applicable
41. Coxal apodemes I
0) free medially Fig-ch41-0.jpg Map-MedeusV.gif?63,17,100,107
1) fused medially with coxal apodemes II closing coxal fields I Fig-ch41-1.jpg Map-MedeusV.gif?53,42,118,112
2) non-applicable because coxae free
42. Number of setae on tibiae I-II (excluding solenidia)
0) 2 Map-DiadasiopusV.htm?75,33,30,20
1) 1 Map-DiadasiopusV.htm?75,33,30,20
2) 3 or more
3) 0
43. Setae g of male
0) filiform Map-MedeusV.gif?96,153,27,26
1) absent Map-MedeusV.gif?96,153,27,26
44. Setae 4a of male
0) filiform Map-MedeusV.gif?85,160,51,41
1) absent/non-applicable Map-MedeusV.gif?85,160,51,41
45. Pretarsal empodium II
0) claw-like Map-DiadasiopusD.htm?187,51,26,26
1) claw-like, rayed with tenent hairs
2) membranous with setules Fig-ch45-2.jpg
3) membranous, nude Fig-ch45-3.jpg
4) absent
46. Prodorsal trichobothria
0) absent
1) 1 pair Fig-ch46-1.jpg
2) 2 pairs Fig-ch46-2.jpg
47. Stigmatal openings
0) absent
1) present, located at or on cheliceral bases
2) present, anterio-lateral on propodosoma Fig-ch47-2.jpg
3) present, situated on lateral sides of body Fig-ch47-3.jpg
48. Peritremes
0) absent
1) present, chambered, arch-like, associated with bases of chelicerae
2) present, usually linear, situated on lateral sides of body; entering stigma anteriorly Fig-ch48-2.jpg
3) present, linear, situated on lateral sides of body; entering stigma posteriorly Fig-ch48-3.jpg
49. Enlarged dorsal claw-like seta on palp tibia
0) absent Fig-ch49-0.jpg
1) present Fig-ch49-1.jpg
2) palps absent or reduced, with 1-3 free non-articulating palopmeres (small claw-like seta may present) Fig-ch49-2.jpg
50. Palp tarsus or, if reduced, tarsal solenidion
0) terminal
1) displaced ventrally
2) not-applicable, because palp absent
51. Solenidia on tibia
0) not recessed into pit
1) at least one recessed into pit Fig-ch51-1.jpg
2) absent, or imposible to determine because podomeres fused
52. Dorsal opisthosomal trichobothria
0) absent Fig-ch52-0.jpg
1) present Fig-ch52-1.jpg
53. Palp
0) with 3 free palpomeres, extending to or slightly beyond subcapitular extremity
1) with 4-5 articulated free palpomeres, extending beyond subcapitular extremity
2) very small, with 1-2 non-articulating free palpomeres, not extending to or slightly beyond subcapitular extremity
3) very small, with 1-2 non-articulating free palpomeres, distinctly extending beyond subcapitular extremity (if present)
4) absent
54. Comb-like setae on palp tarsus (not to be confused with bi- or trifid apotele)
0) absent or palp absent Fig-ch54-0.jpg
1) present, 1-2 pairs Fig-ch54-1.jpg
55. Larval Claparede's organ
0) absent Fig-ch55-0.jpg
1) present Fig-ch55-1.jpg
56. Postero-apical corner of coxa III
0) without reticulate pattern
1) with distinct reticulate pattern Fig-ch56-1.jpg
2) not-applicable because coxa III absent Fig-ch56-2.jpg
57. Tarsus II setation
0) 0 setae
1) 8 setae
2) 1-8 setae
3) more than 8 setae
58. Hysterosomal shield of female
0) divided transversely, anterior part entire
1) divided transversely, anterior part also divided into medial and 2 lateral plates
2) entire (sometimes fused with other shields)
3) absent
59. Tibia and tarsus I of female
0) free
1) fused
60. Pair of internal sacs (sporothecae) between legs III and IV in female
0) absent
1) present
61. Genital capsule of male
0) absent, not icorporating genital and aggenital setae
1) present, incorporating genital and aggenital setae
62. Pharyngeal musculature in female
0) not subdivided or indistinct, or not as on Fig.
1) subdivided into large anterior part and 1 or 2 separate posterior parts (Fig.) Fig-ch62-1.jpg
63. Leg IV of female
0) with separate femur and genu, often pretarsus with claws and membranous empodium
1) with fused femur and genu, pretarsus without claw and empodium
2) absent
64. Leg IV of male
0) with 5 or more podomeres
1) with 3-4 podomeres
2) absent
65. Legs IV of female
0) with 4-5 or more podomeres, not attenuate
1) with 3 podomeres, attenuate
2) absent
66. Movable digit of female chelicera (not to be confused with excrescenses that also may be present at base of digit)
0) with several pectinate, dendritic, brush-like or filamentous excrescenses
1) without excrescenses
67. Anterior margin of epigynial shield of female
0) shield absent, if present (=mesogynial shield) <u>then</u> flanked or covered by latigynial shields
1) rounded or truncate (if pointed than shield weakly sclerotized)
2) triangular, pointed, well-sclerotized
68. latigynial shields in female
0) absent
1) present, mesogynial shield more than 1/2 of length of latigynial shields
2) present, mesogynial shield less than 1/3 of length of latigynial shields
69. Spermatodactyl in male
0) absent
1) present, distal end free Fig-ch69-1.jpg
2) present, distal end fused with movable digit (spermatotreme) Fig-ch69-2.jpg
3) chelicera absent
70. Distinctive spurs, enlarged setae or flanges on male legs
0) absent
1) present
71. Pretarsus I in female and immatures
0) without claws or empodium/pulvillus
1) with claw(s) or empodium/pulvillus Fig-ch71-1.jpg
72. Unpaired postanal seta
0) absent
1) present
73. Anal shield of female
0) oval, not-triangular
1) triangular
2) fused with ventral shield
3) absent
74. Ventromarginal shield of female (not to be confused with metapodal shield)
0) not fused with peritrematal plate
1) fused with peritrematal plate
2) absent
75. Corniculi
0) thin, almost filiform
1) strong, spiniform; not divided distally
2) strong, spiniform; divided distally
3) absent or gnathosoma vestigial or absent
76. Foveolae pedales
0) absent
1) present
77. Pilus dentilis on fixed digit of chelicera
0) simple, absent, or chelicera absent
1) bifurcated
78. Metasternal shields
0) not flanking epigynial shield (if present), rounded, with convex inner edges, or absent
1) flanking epigynial shield, with straight inner edges
79. Palpal apotele
0) absent, or palps absent
1) trifid Fig-ch79-1.jpg
2) bifid Fig-ch79-2.jpg
80. One or two brush-like arthropodial processes at base of movable digit of chelicera
0) absent, or chelicera absent Fig-ch80-0.jpg
1) present Fig-ch80-1.jpg
81. Dorsal idiosoma with
0) fewer than 21 pairs of setae
1) more than 21 pairs of setae
82. Posterior margin of epigynial shield of female
0) shield absent, if present (=mesogynial shield) <u>then</u> flanked or covered by latigynial shields <u>or</u> fused with ventral or ventrianal shield
1) truncate or weakly convex (if rounded then anal shield not triangular and ventrianal shield present)
2) rounded or pointed (if truncate then anal shield triangular, or ventrianal shield absent) Fig-ch82-2.jpg