Uenoidae Iwata, 1927

Introduction

This family was previously placed in the family Limnephilidae (Neophylacinae), but later work by Wiggins and others argued effectively for family elevation. Uenoidea comprises 7 genera of which 5 are found in the Nearctic, and 2 in MIchigan. Oligophleboides is principally a western taxa of cool, rapidly flowing streams, and has only rarely been recorded from northern Michigan. Neophylax, on the other hand, is widespread and common in cold to cool lotic waters.

Adults (adapted from Schmid 1998, Vineyard 1990)

1a	a. In males, inferior appendages not prominent, deeply embedded in Ab9	Oligophlebodes minutus (Banks)
	b. In males, armature of Ab10 composed of 2 membranous lobes and 2 ear-shaped appendages
	c. In females, ventral lobes of Ab9 sternite fused together ventral to the vulvar scale, which is only slightly sclerotized
1b	a. In males, inferior appendages more or less prominent, not embedded in Ab9	Neophylax, 2
	b. In males, armature of Ab10 composed to 2-3 pairs of appendages
	c. In females, ventral lobes of Ab9 sternite fused with the lateral margins of the vulvar scale
2a(1b)	Maxillary palpi 3-sgemented (males)	3
2b	Maxillary palpi 5-segmented (females)	6
3a(2a)	Forewing dark brown or black, without white spots also: Tibial spurs 1,2,2	Neophylax fuscus Banks
3b	Forewing reddish brown, with a tan triangle or double-triangle pattern	4
	also: Metatibia with a medial apical spur < 1/3x length of tarsus, thickened basally with a promientn sclerotized shield; In lateral view, inferior appendage with rounded or pointed lateral lobe oriented caudally or ventrally, oriented horizontally or transversely in caudal view; In ventral view, Ab9 with posterior margin of sternum flat or convex
4a(3b)	Hindwing with R2+3 and R4+5 arising separately from R1	Neophylax oligius Ross
4b	Hindwing with R2+3 fused from R1 to the base of the discoidal cell	5
5a(4b)	a. Ab10 with the internal branch separated caudally from the body of segment by less than 1/4 its length	Neophylax aniqua Ross
	b. Ab10 with the external branch large in relation to the internal branch, in caudal view not produced as a free point, and extended ventrally only to the ventral margin of body of Ab10
5b	a. Ab10 with internal branch separated caudally from body of segment by more than 1/4 its length	Neophylax concinnus McLachlan
	b. Ab10 with the external branch small in relation to the internal branch
	also: In ventral view, Ab9 sternum with its posterior margin convex or flat, in ventral view, not excavated laterally and its caudal margin evenly rounded or straight; In lateral view, inferior appendage with the lateral lobe extended caudad as far as the internal branch of Ab10; In dorsal and ventral aspects, lateral lobe of inferior appendage with a prominent median hook
6a(2b)	Forewing dark brown, without white spots	Neophylax fuscus Banks
6b	Forewing reddish brown, with a light brown triangle or double-triangle pattern	7
	also: Lateral vulvar lobes never as heavily sclerotized points
7a(6b)	Lateral vulval lobes extended caudad farther than the median lobe, rounded laterally, terminating in broad triangular points	Neophylax concinnus McLachlan
	also: Ab8 sternum with its posterior margin fused with Ab9 sternum
7b	Lateral vulval lobes shorter than the median lobe	8
8a(7b)	Lateral vulval lobes larger than the median lobes, rounded laterally	Neophylax aniqua Ross
	also: Median vulval lobe produced as a pair of darkly sclerotized points; Apex of Ab10 with a wide V-shaped notch
8b	Lateral vulval lobes reduced, much smaller than the rounded and apically bifurcate median lobes, and produced as slightly convex sclerotized processes	Neophylax oligius Ross
	also: Ab8 sternum narrowest posteriorly; Median vulval lobes with a shallow apical notch, with a conspicuous basomedian sclerotized point

Larvae (adapted from Vineyard 1990, Wiggins 1996)

1a	a. Pronotum with prominent longitudinal ridges	Oligophlebodes minutus (Banks)
	b. Anteromedian notch of mesonotum weakly represented
	c. Case of rock fragments, strongly tapered and cuved, outline smooth
1b	a. Pronotum lacking longitudinal ridges	Neophylax, 2
	b. Anteromedian notch of mesonotum prominent
	c. Case of rock fragments, with larger balast stones along each side
2a	Ab1 with a ventral gill	Neophylax oligius Ross
	also: Pronotum with short and needle-like spines along the anterior margin; Abdomen with dorsal gills present on some segments, posteroventral gills 2x long as other gills; Head usually dark, with a well-defined pale central stripe
2b	Ab1 without a ventral gill	3
3a(2b)	a. Pronotum with spines along the anterior margin long, prominent and blade-like	Neophylax fuscus Banks
	b. Frontoclypeus without a tubercle
	also: Mesonotal scleroites appearing contiguous mesally; Metanotal sa1 sclerite present; Lateral gills present
3b	a. Pronotum with spines along the anterior margin small and fine	4
	b. Frontoclypeus usually with a prominent median elevation or tubercle
4a(3b)	a. Frontoclypeus depressed, with a long peg-like tubercle at least 2x high as its basal width	Neophylax aniqua Ross
	b. Head usually dark
4b	a. Frontoclypeus usually with only a median elevation, occasionally a very short tubercle present	Neophylax concinnus McLachlan
	b. Head light brown

References

Vineyar RN. 1990. Systematics of the caddisfly genus Neophylax McLachlan (Trichoptera: Uenoidae). Ph.D dissertation, University of Toronto. 252 p.
Wiggins GB. 1996. Larvae of the North American caddisfly genera (Trichoptera), 2nd Edition. University of Toronto Press, Toronto, Ontario. xiii + 457 p.

Page created: November 04, 2003 (EB) - Last updated: July 3, 2007 (EB)