Ceratopogonidae - No-See-Ums or Biting Midges

Below is a preliminary list of species based on records in the UMMZ-Insect Division collection (identified by Wirth, Stone and Steyskal) as well as literature sources, principally those of Wirth, Borkent and Grogan.

Alluaudomyia Kieffer, 1913 (Subfamily Ceratopogoninae)

Alluaudomyia bella (Coquillett), 1902 Ceratopogon - widespread, WI
  • Isoecacta poeyi Garrett, 1925
  • Alluaudomyia splendida, authors, not Winnertz
Alluaudomyia megaparamera Williams, 1957
Alluaudomyia needhami Thomsen, 1935 - widespread, WI
Alluaudomyia wirthi Williams, 1957

Atrichopogon Kieffer, 1906 (Subfamily Forcipomyiinae)

Atrichopogon fusculus (Coquillett), 1901 Ceratopogon - widespread
Atrichopogon fusinervis (Malloch), 1915 Ceratopogon - IL
Atrichopogon levis (Coquillett), 1901 Ceratopogon - UMMZ
  • Ceratopogon exilis Coquillett, 1902
Atrichopogon minutus (Meigen), 1830 Ceratopogon
Atrichopogon peregrinus (Johannsen), 1908 Ceratopogon - UMMZ
Atrichopogon websteri (Coquillett), 1901 Ceratopogon - widespread

Bezzia Kieffer, 1899 (Subfamily Ceratopogoninae)

Bezzia (Bezzia) albidorsata Malloch, 1915 - IL
Bezzia (Bezzia) apicata Malloch, 1914 - IL
Bezzia (Bezzia) bivittata (Coquillett), 1905 Ceratopogon - Wirth and Grogan 1983
Bezzia (Bezzia) copiosa (Thomsen), 1935 Probezzia
Bezzia (Bezzia) cockerelli Malloch, 1915 - WI
Bezzia (Bezzia) dentata Malloch 1914 - IL
Bezzia (Pseudobezzia) expolitus (Coquillett), 1901 Ceratopogon - widespread
Bezzia (Pseudobezzia) flavitarsis Malloch, 1914
Bezzia (Bezzia) gibberella Wirth and Grogan, 1983 - Wirth and Grogan 1983
Bezzia (Bezzia) glabra (Coquillett), 1902 Ceratopogon - UMMZ
Bezzia (Bezzia) incerta (Malloch), 1915 Probezzia - IL; synonomyzed by Grogan and Wirth 1983?
Bezzia (Bezzia) opaca (Loew), 1861 Ceratopogon - IL, ON WI
  • Probezzia fulvithorax Malloch, 1915
Bezzia (Bezzia) setulosa (Loew), 1861 Ceratopogon - ON WI
Bezzia (Bezzia) vaicolor (Coquillett), 1902 Ceratopogon - WI, widespread

Ceratopogon Meigen, 1803 (Subfamily Ceratopogoninae)

Ceratopogon (Ceratopogon) culicoidithorax Hoffman, 1926 - ON

Clinohelea Kieffer, 1917 (Subfamily Ceratopogoninae)

Clinohelea bimaculata (Loew), 1861 Ceratopogon
Clinohelea curriei (Coquillett), 1905 Ceratopogon
  • Clinohelea nebulosa (Malloch), 1915

Culicoides Latreille, 1809 (Subfamily Ceratopogoninae)

Culicoides (Oecacta) alexanderi Wirth and Hubert, 1962
Culicoides (Oecacta) arboricola Root and Hoffman, 1937 - WI east and south
Culicoides (Diphaomyia) baueri Hoffman, 1925 - WI west and east
Culicoides (Oecacta) bickleyi Wirth and Hubert, 1962 - WI east and south
Culicoides (Oecacta) biguttatus (Coquillett), 1901 Ceratopogon - UMMZ
Culicoides (Avaritia) chiopterus (Meigen), 1830 Ceratopogon - widespread
Culicoides (Beltranmyia) crepuscularis Malloch, 1915 - IL west and east
Culicoides (Oecacta) denticulatuts Wirth and Hubert, 1962 - MN, ON, WI
  • Culicoides unicorlor, authors, not Coquillett
Culicoides (Oecacta) dickei Jones, 1956 - ON, WI
Culicoides (Oecacta) downesi Wirth and Hubert, 1962
Culicoides (Oecacta) flukei Jones, 1956 - WI east
Culicoides (Oecacta) furensoides Williams, 1955
Culicoides (Oecacta) guttipennis (Coquillett), 1901 Ceratopogon - MN, OH
Culicoides (Diphaomyia) haematopotus Malloch, 1915 - IL west and east
Culicoides (Oecacta) jamnbacki Wirth and Hubert, 1962
Culicoides (Oecacta) nanus Root and Hoffman, 1937 - ON, WI
Culicoides (Avaritia) obsoletus (Meigen), 1818 Ceratopogon - widespread
Culicoides (Oecacta) piliferus Root and Hoffman, 1937 - UMMZ
Culicoides (Avaritia) sanguisuga (Coquillett), 1901 Ceratopogon - widespread
  • Simulium nocivum Harris, 1862
Culicoides (Beltranmyia) sphagnumensis Williams, 1955
Culicoides (Oecacta) spinosus Root and Hoffman, 1937 - ON, WI
Culicoides (Oecacta) stellifer (Coquillett), 1901 Ceratopogon - ON west and southeast
Culicoides (Oecacta) stilobezzioides Foote and Pratt, 1954 - MN east
Culicoides (Oecacta) travisi Vargas, 1949 - ON, WI
  • Culicoides simulans Root and Hoffman, 1937
  • Culicoides horneae Foote and Pratt, 1954
Culicoides (Monoculicoides) variipennis (Coquillett), 1901 Ceratopogon - widespread
Culicoides (Hoffmania) venustus Hoffman, 1925 - widespread east
Culicoides (Oecacta) villosipennis Root and Hoffman, 1937 - WI southwest and east
Culicoides (Beltranmyia) wisconsinensis Jones, 1956

Dasyhelea Kieffer, 1911 (Subfamily Dasyheleinae)

Dasyhelea grisea (Coquillett), 1901 Ceratopogon
Dasyhelea major (Malloch), 1915 Pseudoculicoides
Dasyhelea mutabilis (Coquillett), 1901 Ceratopogon - widespread
Dasyhelea oppressa Thomsen, 1935 - IA east
Dasyhelea traverae Thomsen, 1935

Echinohelea Macfie, 1940 (Subfamily Ceratopogoninae)

Echinohelea lanei Wirth 1951

Forcipomyia Meigen, 1818 (Subfamily Forcipomyiinae)

Forcipomyia (Forcipomyia) aurea Malloch, 1915 - IL
Forcipomyia (Forcipomyia) bipunctata (Linnaeus), 1767 Tipula - widespread
Forcipomyia (Forcipomyia) bystraki Grogan and Wirth, 1975 - Grogan and Wirth 1975
Forcipomyia (Forcipomyia) cilipes (Coquillett), 1900 Ceratopogon - widespread
Forcipomyia (Forcipomyia) concolor Malloch, 1915 - IL
Forcipomyia (Forcipomyia) elegantula Malloch, 1915 - IL, widespread
Forcipomyia (Neoforcipomyia) eques (Johannsen), 1908 Ceratopogon
Forcipomyia (Lasiohelea) fairfaxensis Wirth, 1951
Forcipomyia (Forcipomyia) fuliginosa (Meigen), 1818 Ceratopogon - NAm widespread
  • Forcipomyia crucicida Knab, 1914
  • Ceratopogon coquilletti Kieffer, 1917
  • Focipomyia brookmani Wirth, 1952
Forcipomyia (Euforcipomyia) hirtipennis (Malloch), 1915 Ceratopogon - IL
Forcipomyia (Neoforcipomyia) mcateei Wirth, 1956
Forcipomyia (Thyridomyia) monilicornis (Coquillett), 1905 Ceratopogon - widespread
Forcipomyia (Thyridomyia) monilicornis (Coquillett), 1905 Ceratopogon - widespread
Forcipomyia (Metaforcipomyia) pluvialis Malloch, 1923 - ON south
Forcipomyia (Caloforcipomyia) splendida Wirth, 1951
Forcipomyia (Forcipomyia) texana (Long), 1902 Ceratopogon - widespread

Heteromyia Say, 1825 (Subfamily Ceratopogoninae)

Heteromyia prattii Coquillett, 1902 - Wirth and Grogan, 1977

Jenkinshelea Macfie, 1934 (Subfamily Ceratopogoninae)

Jenkinshelea albaria (Coquillett), 1895 Ceratopogon - IL, ON, south and east
Jenkinshelea magnipennis (Johannsen), 1908 Johannseniella - Grogan and Wirth 1977

Johannsenomyia Malloch, 1915 (Subfamily Ceratopogoninae)

Johannsenomyia annulicornis Malloch, 1918
Johannsenomyia argentata (Loew), 1861 Ceratopogon - widespread

Leptoconops Skuse, 1889 (Subfamily Leptoconopinae)

Leptoconops (Holoconops) catawbae (Boesel), 1948 Holoconops

Mallochohelea Wirth, 1962 (Subfamily Ceratopogoninae)

Mallochohelea albibasis (Malloch), 1915 Johannsenomyia - IL east, west and south
Mallochohelea albihalter Wirth, 1962
Mallochohelea atripes Wirth, 1962
Mallochohelea flavidula (Malloch), 1914 Johannsenomyia - IL, WI
Mallochohelea smithi (Lewis), 1956 Johannsenomyia - WI east and south
Mallochohelea variegata Wirth, 1962

Monohelea Kieffer, 1917 (Subfamily Ceratopogoninae)

Monohelea (Monohelea) johannseni Wirth, 1953
Monohelea (Monohelea) macflei Wirth, 1953
Monohelea (Monohelea) stonei Wirth, 1953 - ON, south
Monohelea (Schizohelea) leucopeza (Meigen), 1804 - widespread
  • Ceratopogon politus Colquillett, 1901

Nilobezzia Kieffer, 1921 (Subfamily Ceratopogoninae)

Nilobezzia mallochi Wirth, 1962
Nilobezzia minor (Wirth), 1952 Sphaeromias - widespread north

Palpomyia Meigen, 1818 (Subfamily Ceratopogoninae)

Palpomyia altispina Grogan and Wirth 1975 - WI
Palpomyia basalis (Walker), 1848 Ceratopogon - Grogan and Wirth 1979
Palpomyia canadensis Grogan and Wirth 1979 - widespread
Palpomyia downesi Grogan and Wirth 1979 - widespread north
Palpomyia flaviceps (Meigen), 1804 Ceratopogon - Grogan and Wirth 1979
  • Ceratopogon basalis Walker, 1848
  • Johannseniella flaviceps Johannsen, 1908
Palpomyia illinoisensis Malloch, 1914 - IL east; synonomized by Grogan and Wirth 1975?
Palpomyia jonesi Grogan and Wirth 1975 - Grogan and Wirth 1979
Palpomyia lineata (Meigen), 1818 Ceratopogon - Grogan and Wirth 1979
Palpomyia novitibialis Grogan and Wirth 1975 - Grogan and Wirth 1979
Palpomyia opacithorax (Malloch), 1915 Heteromyia - IL east; synonomized by Grogan and Wirth 1975?
Palpomyia plebeiella Grogan and Wirth 1975 - Grogan and Wirth 1979
Palpomyia plebeia (Loew), 1861 Ceratopogon
Palpomyia praeustus (Loew), 1869 Ceratopogon
Palpomyia rubiginosa Grogan and Wirth 1975 - Grogan and Wirth 1979
Palpomyia rufa (Loew), 1861 Ceratopogon - Grogan and Wirth 1979
Palpomyia scalpellifera Grogan and Wirth 1975 - Grogan and Wirth 1979
Palpomyia slossonae (Coquillett), 1905 Ceratopogon - widespread; synonomyzed by Grogan and Wirth 1975?
Palpomyia subaspera (Coquillett), 1901 Ceratopogon - Grogan and Wirth 1979
Palpomyia tenuicornis (Malloch), 1915 Heteromyia - WI; synonomyzed by Grogan and Wirth 1975?
Palpomyia tibialis (Meigen), 1818, Ceratopogon
Palpomyia trivialis (Loew), 1861, Ceratopogon (synonomyzed by Grogan and Wirth?)

Parabezzia Malloch, 1915 (Subfamily Ceratopogoninae)

Parabezzia petiolata Malloch, 1915 - IL
Parabezzia williamsi Wirth 1965 - Grogan and Wirth 1977

Probezzia Kieffer, 1906 (Subfamily Ceratopogoninae)

Probezzia albiventris (Loew), 1861 Ceratopogon - WI south and east
Probezzia atriventris Wirth, 1951
Probezzia concinna (Meigen), 1818 Ceratopogon - ON west
Probezzia infuscata Malloch, 1915 - IL, MN, ON
Probezzia pallida Malloch, 1914 - IL, ON, WI
Probezzia sabroskyi Wirth, 1951
Probezzia smithii (Coquillett), 1901 Ceratopogon - WI east and south
Probezzia concinna (Meigen), 1818 Ceratopogon - ON west
Probezzia williamsi Wirth 1971 - WI
Probezzia xanthogaster (Kieffer), 1917 Bezzia - IL, ON, WI
  • Probezzia elegans Colquillett, 1901

Serromyia Meigen, 1818 (Subfamily Ceratopogoninae)

Serromyia borealis Borkent, 1990
Serromyia crassifermorata Malloch, 1914
Serromyia ledicola Kieffer 1925 - ON, widespread
Serromyia nudicolis Borkent 1990
Serromyia vockerothi Borkent 1990 - MN, widespread north

Sphaeromias Curtis, 1829 (Subfamily Ceratopogoninae)

Sphaeromias longipennis (Loew), 1861 Ceratopogon - MN, ON, WI

Stilobezzia Kieffer, 1911 (Subfamily Ceratopogoninae)

Stilobezzia (Stilobezzia) antennalis (Coquillett), 1901 Ceratopogon - widespread north, ON
Stilobezzia (Stilobezzia) coquilletti Kieffer, 1917 - IL
Stilobezzia (Stilobezzia) sybleae Wirth, 1953
Stilobezzia (Neostilobezzia) lutea (Malloch), 1918 Hartomyia
  • Stilobezzia mallochi, Hoffman, 1924

Page created: November 23, 2002 - Last updated: April 2, 2006 (EB)